Pseudoscorpions of the World

Last updated: 19 January 2009

Version 1.2, 2009

Family CHEIRIDIIDAE Hansen

Cheiridiidae Hansen: Chamberlin, 1931a: 236; Beier, 1932e: 3; Chamberlin, 1935b: 480; Roewer, 1937: 274-275; Chamberlin, 1938a: 264; Petrunkevitch, 1955: 82; Hoff, 1956c: 3; Morikawa, 1960: 133-134; Dubinin, 1962: 442; Meinertz, 1962: 60; Vitali-di Castri, 1962: 120; Beier, 1963b: 243; Vitali-di Castri, 1965: 69; Murthy and Ananthakrishnan, 1977: 115; Harvey, 1985b: 130; Muchmore, 1982a: 99-100; Legg and Jones, 1988: 90; Harvey, 1991a: 433; Harvey, 1992c: 1416-1417.

Subfamily CHEIRIDIINAE Hansen

Cheiridiinae Hansen, 1894: 232 (as Chiridiinae); With, 1906: 116 (as Chiridiinae); Lessert, 1911: 924 (as Chiridiinae); Chamberlin, 1924a: 33-34; Väänänen, 1928a: 17 (as Chiridiinae); Beier, 1932e: 6-7; Beier, 1932g: 185; Roewer, 1937: 276; Morikawa, 1960: 134; Meinertz, 1962: 60; Murthy and Ananthakrishnan, 1977: 115-116.

Genus Apocheiridium Chamberlin

Apocheiridium Chamberlin, 1924a: 34; Chamberlin, 1931a: 238; Beier, 1932e: 10; Chamberlin, 1935b: 480; Hoff, 1952b: 192; Morikawa, 1960: 135; Beier, 1963b: 244-245; Beier, 1976f: 211; Murthy and Ananthakrishnan, 1977: 116; Benedict, 1978b: 232; Harvey, 1991a: 433.

 

Type species: Apocheiridium ferumoides Chamberlin, 1924a, by original designation.

Distribution: AFRICA: widespread.

ASIA: widespread.

AUSTRALASIA: widespread.

EUROPE: widespread.

NORTH AMERICA: widespread.

SOUTH AMERICA: Chile.

 

Subgenus Apocheiridium (Apocheiridium) Chamberlin

Apocheiridium (Apocheiridium) Chamberlin: Harvey, 1991a: 433.

Apocheiridium (Apocheiridium) asperum Beier

Apocheiridium asperum Beier, 1964k: 68-69, fig. 27.

Apocheiridium (Apocheiridium) asperum Beier: Harvey, 1991a: 433.

 

Type locality: Masite, Lesotho.

Distribution: Lesotho.

Apocheiridium (Apocheiridium) bulbifemorum Benedict

Apocheiridium bulbifemorum Benedict, 1978b: 233-234, fig. 1.

Apocheiridium (Apocheiridium) bulbifemorum Benedict: Harvey, 1991a: 433.

 

Type locality: 13 km S., 6 km E. of Tiller, Douglas County, Oregon, U.S.A.

Distribution: U.S.A. (Oregon).

Apocheiridium (Apocheiridium) caribicum Beier

Apocheiridium caribicum Beier, 1936a: 446, fig. 3; Roewer, 1940: 346; Muchmore, 1993a: 92.

Apocheiridium (Apocheiridium) caribicum Beier: Harvey, 1991a: 433.

 

Type locality: Bak Ariba, Hato, Curaçao, Netherlands Antilles.

Distribution: Netherlands Antilles (Curaçao).

Apocheiridium (Apocheiridium) cavicola Mahnert

Apocheiridium cavicola Mahnert, 1993b: 989-992, figs 43-47; Ashmole and Ashmole, 1997: 556, 579.

 

Type locality: Ravin Cave, Ascension Island, St Helena.

Distribution: St Helena (Ascension Island).

Apocheiridium (Apocheiridium) chamberlini Godfrey

Cheiridium ferum Simon: Ellingsen, 1912b: 86, 104 (misidentification); Godfrey, 1923: 95-98 (misidentification, in part).

Apocheiridium sp.: Chamberlin, 1924a: 37.

Apocheiridium chamberlini Godfrey, 1927: 17-18; Chamberlin, 1932b: 139; Beier, 1932e: 13; Roewer, 1937: 277; Beier, 1964k: 67-68, fig. 26.

Apocheiridium (Apocheiridium) chamberlini Godfrey: Harvey, 1991a: 434.

 

Type locality: Lovedale, Eastern Cape, South Africa.

Distribution: South Africa.

Apocheiridium (Apocheiridium) eruditum Chamberlin

Apocheiridium eruditum Chamberlin, 1932b: 139; Beier, 1932e: 13; Roewer, 1937: 277.

Apocheiridium (Apocheiridium) eruditum Chamberlin: Harvey, 1991a: 434.

 

Type locality: Los Baños, Luzon Island, Philippines.

Distribution: Philippines.

Apocheiridium (Apocheiridium) fergusoni Benedict

Apocheiridium fergusoni Benedict, 1978b: 234-235, fig. 2; D.W. Zeh, 1987b: 1087.

Apocheiridium (Apocheiridium) fergusoni Benedict: Harvey, 1991a: 434.

 

Type locality: 18 km SE. of Riley, Harney County, Oregon, U.S.A.

Distribution: U.S.A. (Oregon).

Apocheiridium (Apocheiridium) ferum (Simon)

Cheiridium ferum Simon, 1879a: 44-45, plate 18 fig. 21 (as Chiridium [sic] ferum); Ellingsen, 1907a: 166-167; With, 1907: 80-81, figs 54-57 (as Chiridium [sic] ferum); Warburton, 1909: 437 (as Chiridium [sic] ferum); Lessert, 1911: 25-26, fig. 20 (as Chiridium [sic] ferum); Kew, 1914: 96-97; Godfrey, 1923: 95-98 (in part; see Apocheiridium (Apocheiridium) chamberlini Godfrey); Redikorzev, 1924b: 25, fig. 11a; Kästner, 1927a: 15; Caporiacco, 1928b: 127 (as Chiridium [sic] ferum); Kästner, 1928: 4, fig. 11.

Apocheiridium ferum (Simon): Chamberlin, 1924a: 37; Beier, 1929a: 358; Chamberlin, 1931a: fig. 29n; Beier, 1932e: 11-12, fig. 7; Chamberlin, 1934b: 7; Redikorzev, 1935a: fig. 2; Roewer, 1937: 277, fig. 224; Vachon, 1952c: 537-539; Beier, 1956h: 25; Beier, 1956i: 8; Ressl and Beier, 1958: 2; Beier, 1963b: 245, fig. 249; Dumitresco and Orghidan, 1964: 624; Orghidan and Dumitresco, 1964a: 328; Orghidan and Dumitresco, 1964b: 192; Beier, 1965f: 93; Ressl, 1965: 289; Dumitresco and Orghidan, 1966: 81-82; Helversen, 1966a: 138-139, fig. 2; Weygoldt, 1966d: 14, 15, 22; Zangheri, 1966: 531; Rafalski, 1967: 14-15; Decou and Negrea, 1969: 523; Lazzeroni, 1969c: 240; Weygoldt, 1969a: 26, 41, 74, fig. 51; Verner, 1971: fig. 46; Callaini, 1979c: 350; Schaller, 1979: plate 10.III fig. 6; Thaler, 1979: 52; Callaini, 1982a: 17, 23, fig. 4c; Callaini, 1983a: 152-153; Schaefer, 1988: 99; Schawaller and Dashdamirov, 1988: 34, figs 56-57; Pieper, 1991: 218, 220; Dashdamirov and Schawaller, 1992b: 54, fig. 11; Dashdamirov and Schawaller, 1993b: 2, figs 1-2; Gardini, 1995a: 6; Platen, Blick, Bliss, Drogla, Malten, Martens, Sacher and Wunderlich, 1995: 54; Mahnert, 1997: 570; Dashdamirov, 1999: 82; Ducháč, 1999a: 14; Gardini, 2000: 107; Kovoor and Muñoz-Cuevas, 2000: 45, 64; Petrov, 2000: 62-63, fig. 1; Petrov, 2001: 70; Šťáhlavský and Ducháč, 2001: 48; Delfosse, 2003: 38; Drogla and Lippold, 2004: 26; Mahnert, 2004: 464.

Cheiridium aff. ferum Simon: Protescu, 1937: 97-99, figs 12-15.

Apocheiridium (Apocheiridium) ferum (Simon): Harvey, 1991a: 434; Georgescu and Capuse, 1996a: 111.

Not Cheiridium ferum Simon: Ellingsen, 1912b: 86, 104 (misidentification; see Apocheiridium (Apocheiridium) chamberlini Godfrey).

 

Type locality: Arcachon, Gironde, Aquitaine, France.

Distribution: Austria; Azerbaijan; Bulgaria; Czech Republic; Estonia; France (mainland); Germany; Italy (mainland, Sardinia); Poland; Portugal (Madeira); Romania; Samoa (extremely doubtful); Spain (Islas Canarias); Switzerland; Turkey; Uzbekistan.

Apocheiridium (Apocheiridium) ferumoides Chamberlin

Apocheiridium ferumoides Chamberlin, 1924a: 35-36, figs b-f, h, k-p, r-s, v, x-y, aa (in part, see Apocheiridium (Apocheiridium) inexpectum Chamberlin); Beier, 1930a: 207; Chamberlin, 1931a: figs 6g, 14g, 15t, 18c-d, 26f, 38e-g, 44h-i, 47u, 51c-e, 52h, 66; Chamberlin, 1932b: 139; Beier, 1932e: 12, figs 8-9; Beier, 1932g: fig. 261; Vachon, 1934i: 414 (as Apocheiridium fermuroides [sic]); Chamberlin, 1935b: 480; Roewer, 1936: figs 20, 43c-d; Roewer, 1937: 277; Vachon, 1949: figs 200a, 203g (as Apocheiridium femuroides [sic]); Gering, 1956: 50; Essig, 1958: 11, figs 7a-c; Hoff, 1958: 19; Benedict, 1978b: 235-237; D.W. Zeh, 1987b: 1087; Judson, 1997a: 17.

Apocheiridium (Apocheiridium) femuroides Chamberlin: Harvey, 1991a: 434.

 

Type locality: Stanford University, Palo Alto, Santa Clara County, California, U.S.A.

Distribution: U.S.A. (California, Oregon, Utah).

Apocheiridium (Apocheiridium) granochelum Benedict

Apocheiridium granochelum Benedict, 1978b: 237-238, figs 3-5; D.W. Zeh, 1987b: 1087.

Apocheiridium (Apocheiridium) granochelum Benedict: Harvey, 1991a: 435.

 

Type locality: Corvallis, Benton County, Oregon, U.S.A.

Distribution: U.S.A. (Oregon).

Apocheiridium (Apocheiridium) indicum Murthy and Ananthakrishnan

Apocheiridium indicum Murthy and Ananthakrishnan, 1977: 116-117, fig. 38; Bhattacharyya, 1990a: 245, fig. 1.

Apocheiridium (Apocheiridium) indicum Murthy and Ananthakrishnan: Harvey, 1991a: 435.

 

Type locality: Shoranur, Kerala, India.

Distribution: India.

Apocheiridium (Apocheiridium) inexpectum Chamberlin

Apocheiridium ferumoides Chamberlin, 1924a: 34 (misidentification, in part).

Apocheiridium inexpectum Chamberlin, 1932b: 139-140; Beier, 1932e: 13; Roewer, 1937: 277; Hoff, 1958: 19; Benedict, 1978b: 238-239; D.W. Zeh, 1987b: 1087.

Apocheiridium (Apocheiridium) inexpectum Chamberlin: Harvey, 1991a: 435.

 

Type locality: Beaumont, Riverside County, California, U.S.A.

Distribution: Mexico; U.S.A. (California).

Apocheiridium (Apocheiridium) minutissimum Beier

Apocheiridium minutissimum Beier, 1964d: 312, fig. 1; Tenorio and Muchmore, 1982: 378.

Apocheiridium (Apocheiridium) minutissimum Beier: Harvey, 1991a: 435.

 

Type locality: Subang, Selangor, Malaysia.

Distribution: Malaysia (mainland).

Apocheiridium (Apocheiridium) mormon Chamberlin

Apocheiridium mormon Chamberlin, 1924a: 36-37, figs j, jj, p-q, z; Beier, 1932e: 12-13; Roewer, 1937: 277; Hoff, 1958: 19; Benedict, 1978b: 239-241; D.W. Zeh, 1987b: 1087.

Apocheiridium (Apocheiridium) mormon Chamberlin: Harvey, 1991a: 435.

 

Type locality: Fish Haven, Bear Lake, Bear Lake County, Idaho, U.S.A.

Distribution: U.S.A. (Idaho, Oregon, Utah).

Apocheiridium (Apocheiridium) pallidum Mahnert

Apocheiridium sp.: Mahnert, 1981c: fig. 2.

Apocheiridium pallidum Mahnert, 1982b: 125-127, figs 30-33; Beron, 2002: 36.

Apocheiridium (Apocheiridium) pallidum Mahnert: Harvey, 1991a: 435.

 

Type locality: Morijo, Loita Hills, Rift Valley, Kenya.

Distribution: Kenya.

Apocheiridium (Apocheiridium) pelagicum Redikorzev

Apocheiridium pelagicum Redikorzev, 1938: 87-89, fig. 16; Dawydoff, 1940: 447-449; Roewer, 1940: 346; Vachon, 1940f: 3; Beier, 1948b: 461; Beier, 1951a: 70; Roth and Brown, 1976: 127; Jędryczkowski, 1998: 186.

Apocheiridium (Apocheiridium) pelagicum Redikorzev: Harvey, 1991a: 435.

 

Type locality: Poulo Dama, Cambodia; Cu Lao Cham, Nha Trang (as Nhatrang), Khanh Hoa, Vietnam; and Côn So’n (as Poulo-Condore), Khanh Hoa, Vietnam.

Distribution: Cambodia; Vietnam.

Apocheiridium (Apocheiridium) pinium Morikawa

Apocheiridium pinium Morikawa 1953a: 347-348, figs 1d-g; Morikawa, 1960: 135, plate 3 fig. 5, plate 8 fig. 15, plate 9 fig. 16, plate 10 fig. 20; Morikawa, 1962: 419; Sato, 1978c: 41; Sato, 1979a: 100-102, figs 11a-o, plates 8, 17; Sato, 1979d: 44; Sato, 1980b: 65-71, figs 1-3; Sato, 1980d: plate 6; Sato, 1982c: 34; Sato, 1984d: fig. 2; Sato, Takano and Nakajima, 1988: 24; Sato, 1994b: 80.

Apocheiridium (Apocheiridium) pinium Morikawa: Harvey, 1991a: 435-436.

 

Type locality: Ishite River, Matsuyama, Ehime Prefecture, Shikoku, Japan.

Distribution: Japan.

Apocheiridium (Apocheiridium) reddelli Muchmore

Apocheiridium sp.: Reddell, 1970: 402.

Apocheiridium reddelli Muchmore, 1992: 139-140, figs 16-18.

 

Type locality: Devil's Sinkhole, near Rocksprings, Edwards County, Texas, U.S.A.

Distribution: U.S.A. (Texas).

Apocheiridium (Apocheiridium) rossicum Redikorzev

Apocheiridium rossicum Redikorzev, 1935a: 184-185, figs 1-2; Roewer, 1937: 277; Vachon, 1940f: 3; Beier, 1948b: 461; Kaisila, 1949b: 81, map 5; Dashdamirov and Schawaller, 1993b: 3-5, figs 3-7; Beron, 2002: 36, 39.

Apocheiridium nepalense Ćurčić, 1980b: 80-83, figs 10-14; Schawaller, 1987c: 212; Schawaller, 1991b: 781, figs 1, 18-19; Judson, 1997a: 29 (as synonym of Apocheiridium (Apocheiridium) rossicum Redikorzev) (synonymised by Dashdamirov and Schawaller, 1993b: 3).

Apocheiridium (Apocheiridium) nepalense Ćurčić: Harvey, 1991a: 435.

Apocheiridium (Apocheiridium) rossicum Redikorzev: Harvey, 1991a: 436.

 

Type locality: of Apocheiridium rossicum: Merreküll, Estonia; Sankt-Peterburg (as Petersburg), Leningradskaya Oblast', Russia; and Perm', Permskaya Oblast’, Russia.

of Apocheiridium nepalense: Thini Kola, Nepal.

Distribution: Estonia; Finland; Kyrgyzstan; Nepal; Russia (Leningradskaya Oblast’, Permskaya Oblast’).

Apocheiridium (Apocheiridium) stannardi Hoff

Apocheiridium stannardi Hoff, 1952b: 193-195, figs 3-4; Hoff and Bolsterli, 1956: 164; Hoff, 1958: 19; Hoff, 1961: 444; Nelson, 1971: 95; Nelson, 1975: 283-284, figs 35-39; D.W. Zeh, 1987b: 1087; Snider and Nelson, 1991: 261.

Apocheiridium (Apocheiridium) stannardi Hoff: Harvey, 1991a: 436.

 

Type locality: Herod, Pope County, Illinois, U.S.A.

Distribution: U.S.A. (Colorado, Illinois, Michigan).

Apocheiridium (Apocheiridium) turcicum Beier

Apocheiridium turcicum Beier, 1967f: 308-310, fig. 3.

Apocheiridium (Apocheiridium) turcicum Beier: Harvey, 1991a: 436.

 

Type locality: Namrun, İçel, Turkey.

Distribution: Turkey.

Apocheiridium (Apocheiridium) validissimum Beier

Apocheiridium validissimum Beier, 1976f: 212-213, fig. 13; Marris, 2000: 185.

Apocheiridium (Apocheiridium) validissimum Beier: Harvey, 1991a: 436.

 

Type locality: Mangahou Creek, Chatham Island, New Zealand.

Distribution: New Zealand.

Apocheiridium (Apocheiridium) validum Beier

Apocheiridium validum Beier, 1967b: 292-293, fig. 10; Beier, 1976f: 212-213; Palma, Lovis and Tither, 1989: 20.

Apocheiridium (Apocheiridium) validum Beier: Harvey, 1991a: 436.

 

Type locality: Totara Reserve, Pohangina Valley, Palmerston North, Wellington, North Island, New Zealand.

Distribution: New Zealand.

Apocheiridium (Apocheiridium) zealandicum Beier

Apocheiridium zealandicum Beier, 1976f: 212, fig. 12.

Apocheiridium (Apocheiridium) zealandicum Beier: Harvey, 1991a: 436.

 

Type locality: Stokes Valley, North Island, New Zealand.

Distribution: New Zealand.

Subgenus Apocheiridium (Chiliocheiridium) Vitali-di Castri

Apocheiridium (Chiliocheiridium) Vitali-di Castri, 1969b: 267; Harvey, 1991a: 436.

 

Type species: Apocheiridium leopoldi Vitali-di Castri, 1962, by original designation.

Distribution: SOUTH AMERICA: Chile.

Apocheiridium (Chiliocheiridium) chilense Vitali-di Castri

Apocheiridium chilense Vitali-di Castri, 1962: 123-126, fig. 2; Beier, 1964b: 341; Cekalovic, 1984: 17.

Apocheiridium (Chiliocheiridium) chilense Vitali-di Castri: Vitali-di Castri, 1969b: 267; Harvey, 1991a: 436.

 

Type locality: Quebrada El Tigre, Zapallar, Valparaíso, Chile.

Distribution: Chile.

Apocheiridium (Chiliocheiridium) leopoldi Vitali-di Castri

Apocheiridium leopoldi Vitali-di Castri, 1962: 121-123, fig. 1; Beier, 1964b: 341; Vitali-di Castri, 1966a: fig. 3; Vitali-di Castri, 1966b: figs 7, 9; Cekalovic, 1984: 18.

Apocheiridium (Chiliocheiridium) leopoldi Vitali-di Castri: Vitali-di Castri, 1969b: 267; Harvey, 1991a: 437.

 

Type locality: Cerro El Roble, Valparaíso, Chile.

Distribution: Chile.

Apocheiridium (Chiliocheiridium) serenense Vitali-di Castri

Apocheiridium serenense Vitali-di Castri, 1966b: fig. 8 (nomen nudum).

Apocheiridium (Chiliocheiridium) serenense Vitali-di Castri, 1969b: 267-277, figs 1-25; Cekalovic, 1984: 18; Harvey, 1991a: 437.

 

Type locality: Bahía Guanaqueros, La Serena, Coquimbo, Chile.

Distribution: Chile.

Genus Cheiridium Menge

Cheiridium Menge, 1855: 36; Stecker, 1874b: 232; Stecker, 1875d: 87; Simon, 1879a: 43 (as Chiridium [sic]); Tömösváry, 1882b: 209; Hansen, 1884: 535 (as Chiridium [sic]); Balzan, 1887b: no pagination; Balzan, 1890: 409-410; O.P.-Cambridge, 1892: 229-230; Tullgren, 1899a: 177-178; Planet, 1905: 283-284 (as Chiridium [sic]); Tullgren, 1906a: 201; Kew, 1911a: 50; Lessert, 1911: 24 (as Chiridium [sic]); Chamberlin, 1924a: 37-38; Redikorzev, 1924b: 24; Schenkel, 1928: 56; Chamberlin, 1931a: 237; Beier, 1932e: 7-8; G.O. Evans and Browning, 1954: 14; Chamberlin, 1938a: 264-265; Hoff, 1952b: 188; Morikawa, 1960: 134; Meinertz, 1962: 60-61; Beier, 1963b: 243-244; Legg and Jones, 1988: 91; Harvey, 1991a: 437; International Commission on Zoological Nomenclature, 1992b: 292; Muchmore, 1992: 138.

Cheiridium (Cheiridium) Menge: Chamberlin, 1938a: 265-266.

Cheiridium (Isocheiridium) Chamberlin, 1938a: 269 (synonymised by Hoff and Clawson, 1952: 13-14).

 

Type species: of Cheiridium: Chelifer museorum Leach, 1817, by subsequent designation of Simon, 1879a: 43.

of Cheiridium (Isocheiridium): Cheiridium (Isocheiridium) minor Chamberlin, 1938a, by original designation.

Distribution: AFRICA: widespread.

AMERICAS: widespread.

ASIA: widespread.

EUROPE: widespread.

Cheiridium andinum Vitali-di Castri

Cheiridium andinum Vitali-di Castri, 1962: 126-130, figs 3a-b; Beier, 1964b: 341; Vitali-di Castri, 1969b: 266; Cekalovic, 1984: 18; Harvey, 1991a: 437.

 

Type locality: Hacienda San Vicente, Los Andes, Valparaíso, Chile.

Distribution: Chile.

Cheiridium angustum Beier

Cheiridium angustum Beier, 1978a: 429-430, fig. 1; Harvey, 1991a: 437.

 

Type locality: Uis Tin mine, Okambahe Reserve, Omaruru, Namibia.

Distribution: Namibia.

Cheiridium aokii Sato

Cheiridium aokii Sato, 1984b: 52-55, figs 14-26; Harvey, 1991a: 437; Sato, 1991b: 218, fig. 3.

 

Type locality: Nakano-daira, Haha-jima, Ogasawara-shoto (Bonin Islands), Tokyo Prefecture, Japan.

Distribution: Japan.

Cheiridium brasiliense Mahnert

Cheiridium brasiliense Mahnert, 2001: 117-118, figs 50-54.

 

Type locality: Gruta dos Cristais I, Morro do Chapéu, Bahia, Brazil.

Distribution: Brazil (Bahia).

Cheiridium capense Beier

Cheiridium capense Beier, 1970c: 60-61, fig. 3; Harvey, 1991a: 437.

 

Type locality: Mafeking, North-West Province, South Africa.

Distribution: South Africa.

Cheiridium chamberlini Dumitresco and Orghidan

Cheiridium chamberlini Dumitresco and Orghidan, 1981: 84-86, figs 1c-e, 2c, 4b, 5b, 6b; Armas and Alayón, 1984: 11; Harvey, 1991a: 438.

 

Type locality: Cueva de Peñas Altas de Cativar, El Cobre, Santiago de Cuba, Cuba.

Distribution: Cuba.

Cheiridium congicum Beier

Cheiridium museorum (Leach): Beier, 1955a: 12 (misidentification).

Cheiridium congicum Beier, 1970c: 58-59, fig. 2; Harvey, 1991a: 438.

 

Type locality: Parc National de l'Upemba, Shaba, Democratic Republic of Congo (Zaire).

Distribution: Democratic Republic of Congo (Zaire).

Cheiridium danconai Vitali-di Castri

Cheiridium danconai Vitali-di Castri, 1965: 70-82, figs 1-28; Vitali-di Castri, 1966a: figs 2-3; Vitali-di Castri, 1966b: figs 4, 9; Vitali-di Castri, 1969b: 266; Mahnert, 1981c: fig. 2; Cekalovic, 1984: 18; Harvey, 1991a: 438.

 

Type locality: Monte Amargo, Atacama, Chile.

Distribution: Chile.

Cheiridium fallax Beier

Cheiridium fallax Beier, 1970c: 57-58, fig. 1; Harvey, 1991a: 438.

 

Type locality: Alexandria, Eastern Cape, South Africa.

Distribution: South Africa.

Cheiridium firmum Hoff

Cheiridium firmum Hoff, 1952b: 188-192, figs 1-2; Park and Auerbach, 1954: 210; Hoff, 1958: 18; Gardner, 1986: 22; D.W. Zeh, 1987b: 1087; Harvey, 1991a: 438; Muchmore, 1992: 138-139.

 

Type locality: Pinkstaff, Lawrence County, Illinois, U.S.A.

Distribution: U.S.A. (Illinois, Missouri).

† Cheiridium hartmanni (Menge)

Chelifer hartmanni Menge, in C.L. Koch and Berendt, 1854: 96; Menge, 1855: 38, plate 5 figs 12-13; Scudder, 1891: 255.

Cheiridium hartmanni (Menge): Hagen, 1870: 264; Scudder, 1891: 254; Chamberlin, 1931a: 237; Beier, 1932e: 278; Beier, 1937d: 307, fig. 8; Roewer, 1940: 331; Petrunkevitch, 1955: fig. 50(1); Beier, 1955g: 51; Dubinin, 1962: fig. 1283; Schawaller, 1978: 4; Harvey, 1991a: 438; Selden, 1993b: 305.

Cheiridium sp. ? hartmanni (Menge): Schawaller, 1978: 18-20, figs 11-14.

 

Type locality: Baltic Amber (Eocene).

Distribution: Baltic Amber (Eocene).

Cheiridium insperatum Hoff and Clawson

Cheiridium insperatum Hoff and Clawson, 1952: 8-14, figs 5-8; Hoff, 1958: 18; Muchmore, 1971a: 90; D.W. Zeh, 1987b: 1087; Harvey, 1991a: 438; Muchmore, 1992: 138; Villegas-Guzmán and Pérez, 2005a: 65, 67, 76; Francke and Villegas-Guzmán, 2006: 294.

 

Type locality: Moab, Grand County, Utah, U.S.A.

Distribution: Mexico; U.S.A. (California, Utah).

Cheiridium insulare Vitali-di Castri

Cheiridium insulare Vitali-di Castri, 1984: 1073-1075, figs 38-42; Harvey, 1991a: 438.

 

Type locality: Coreil, Guadeloupe.

Distribution: Guadeloupe.

Cheiridium itapemirinense (Feio)

Geogarypus (Geogarypus) itapemirinensis Feio, 1941: 241-244, fig. 1; Feio, 1945: 5.

Geogarypus itapemirinensis Feio: Harvey, 1986: 760; Harvey, 1991a: 255; Kury and Nogueira, 1999: 10.

Cheiridium itapemirinense (Feio): Mahnert, 2001: 116-117, figs 48-49.

 

Type locality: Cachoeiro de Itapemirim, Espírito Santo, Brazil.

Distribution: Brazil (Distrito Federal, Espírito Santo).

Cheiridium minor Chamberlin

Cheiridium (Isocheiridium) minor Chamberlin, 1938a: 270, figs 2a, 2g, 2i, 2m.

Cheiridium minor Chamberlin: Roewer, 1940: 346; Morikawa, 1953a: 345-347, figs 1a-c; Morikawa, 1960: 134-135, plate 3 fig. 4, plate 10 figs 18, 27; Morikawa, 1962: 419; W.K. Lee, 1981: 129-130, figs 1a-b; Harvey, 1991a: 439; Hong and T.H. Kim, 1993: 2; Sato, 1994b: 80; Schawaller, 1995b: 1056; Dashdamirov, 2005: 245, figs 113-118.

 

Type locality: China.

Distribution: China; Japan; South Korea; Pakistan.

Cheiridium museorum (Leach)

Phalangium cancroides (Linnaeus): Shaw and Nodder, 1791: no pagination (misidentification).

Chelifer nepoides Hermann, 1804: 116, plate 5 fig. q; Théis, 1832: 75-77, plate 3 figs 3a-b; Gervais, 1844: 79; Lucas, 1849: 276; Hagen, 1870: 266; International Commission on Zoological Nomenclature, 1992b: 292 (suppressed for the purposes of the Principle of Priority but not for those of the Principle of Homonymy) (synonymised by Tömösváry, 1882b: 210).

Chelifer museorum Leach, 1817: 50, plate 142 fig. 4; Templeton, 1836: 14 (as Chélifer musaeòrum [sic]); C.L. Koch, 1843: 43-44, fig. 781; Gervais, 1844: 78 (as Chelifer muscorum [sic]); Holmberg, 1876: 28.

Cheiridium museorum (Leach): Menge, 1855: 36-38, plate 1 figs 1, 18, 24, plate 3 figs 4-5, 16-18, plate 4 fig. 11; Hagen, 1870: 264; L. Koch, 1873: 2-3; Pavesi, 1876: 73-74; Pavesi, 1878: 362; Simon, 1879a: 43-44, plate 18 figs 19-20 (as Chiridium [sic] museorum); Becker, 1884: cclxiv (as Chiridium [sic] museorum); Hansen, 1884: 535-536 (as Chiridium [sic] museorum); O.-P. Cambridge, 1886: 45; O.P.-Cambridge, 1892: 230-231, plate C fig. 21 (as Chiridium [sic] museorum); Ritsema, 1874: xxxiii (as Cheiridium musaeorum [sic]); Stecker, 1874b: 232; Stecker, 1875d: 87-88; Pavesi, 1878: 362 (not seen); Simon, 1878a: 152; Daday, 1880a: 192 (as Cheiridium musearum [sic]); Tömösváry, 1882b: 210-211, plate 2 figs 16-18; Canestrini, 1884: no. 7; Daday, 1888: 125, 181; O.-P. Cambridge, 1889: 111; Lameere, 1895: 475; Becker, 1896: 331-332 (as Chiridium [sic] museorum); Pocock, 1896c: 237 (as Chiridium [sic] museorum); Tullgren, 1899a: 178-179, plate 2 figs 1-3; Evans, 1901b: 242 (as Chiridium [sic] museorum); Godfrey, 1901b: 217 (as Chiridium [sic] museorum); Kew, 1901b: 194-195 (as Chiridium [sic] museorum); Kew, 1903: 298; Planet, 1905: 284, unnumbered fig. on p. 283 (as Chiridium [sic] museorum); Tullgren, 1906a: 201, fig. 3; Ellingsen, 1907a: 167; Whyte and Whyte, 1907: 203; Butterfield, 1908: 112; Falconer, 1908: 110 (as Chiridium [sic] museorum); Godfrey, 1908: 158; Jackson, 1908: 73 (as Chiridium [sic] museorum); Godfrey, 1909: 157-159; Kew, 1909a: 249; Kew, 1909b: 259; Warburton, 1909: 437, 438, fig. 226 (as Chiridium [sic] museorum); Ellingsen, 1911c: 174; Kew, 1911a: 51, fig. 13; Lessert, 1911: 24-25 (as Chiridium [sic] museorum); Nonidez, 1917: 26-27 (as Chiridium [sic] museorum); Navás, 1925: 111 (as Chiridium [sic] museorum); Ellingsen, 1912b: 86, 104; Standen, 1912: 14-15; Ellingsen, 1914: 11; Kew, 1914: 96; Falconer, 1916: 191; Kew, 1916a: 124; Kew, 1916b: 79; Standen, 1916b: 124; Standen, 1917: 29; Daday, 1918: 2; Chamberlin, 1924a: 38, figs a, i, t, u, w, bb; Redikorzev, 1924b: 24-25, figs 10, 11b; Kästner, 1927b: figs 10, 41; Kästner, 1928: 4, figs 7, 9-10; Schenkel, 1928: 56, figs 2a-b; Väänänen, 1928a: 17, fig. 6(5) (as Chiridium [sic] museorum); Beier, 1929a: 358; Berland, 1929: 16, fig. (as Chiridium [sic] museorum); Hansen, 1930: plate 15 fig. 2a (as Chiridium [sic] museorum); Chamberlin, 1931a: figs 10b, 15u, 20c, 29k, 33l, 38h; Chamberlin, 1932b: 137; Beier, 1932e: 8-9, figs 2, 5; Tumšs, 1934: 14, fig. 1; Redikorzev, 1935a: fig. 2; Vachon, 1935a: 330-332; Vachon, 1935j: 187; Franganillo, 1936: 155 (misidentification?) (as Chiridium [sic] museorum); Nordberg, 1936: 48-49; Roewer, 1936: figs 43a-b, 70r; Vachon, 1936b: 140; Roewer, 1937: 277; Vachon, 1937f: 128-129; Chamberlin, 1938a: 266-268, figs 2e-f, 2l; Beier, 1939e: 311; Beier, 1939f: 197; Lohmander, 1939b: 292; Morley, 1940: 168 (as Chiridium [sic] musaeorum [sic]); Vachon, 1940a: 53-54; Savory, 1945: 180, 183-184 (as Cheiridium muscorum [sic]); Cooreman, 1946b: 7-8; Beier, 1947b: 323; Kaisila, 1947: 86; Beier, 1948b: 457, 461; Franz and Beier, 1948: 451; Kaisila, 1949b: 80-81, map 5; Redikorzev, 1949: 645-646; Vachon, 1949: fig. 199f; Beier, 1952e: 4; Beier and Franz, 1954: 457; G.O. Evans and Browning, 1954: 14-15, fig. 5; Weidner, 1954b: 109; Beier, 1956h: 24; Cloudsley-Thompson, 1956b: 193; George, 1957: 80; Cloudsley-Thompson, 1958: 96; Ressl and Beier, 1958: 2; Cirdei and Gutu, 1959: 1-2; Cerruti, 1959: 51 (as Cheiridium muscorum [sic]); George, 1961: 38; Meinertz, 1962: 61-62, figs 5d, 36 (as Cheiridium muscorum [sic]); Beier, 1963b: 244, fig. 248; Beier, 1963h: 156; Beier, 1964k: 67; Meinertz, 1964: 392, map 3 (as Cheiridium muscorum [sic]); Weygoldt, 1964a: 362-366, figs 6-11; Ressl, 1965: 289; Weygoldt, 1965a: fig. 47; Weygoldt, 1965c: 436; Beier, 1966k: 464; Helversen, 1966a: 138; Weygoldt, 1966b: fig. 9; Weygoldt, 1966d: 14, 16, 22, 57, fig. 9; Rafalski, 1967: 14; Cloudsley-Thompson, 1968: 126, 127, 139; Kaestner, 1968: 213, 215; Hammen, 1969: 19; Weygoldt, 1969a: 23, 26, 41, 42, 74, 108, 110, figs 23, 38, 51, 89-90, 100; Legg, 1970b: fig. 3(3); Ressl, 1970: 250-251; Helversen and Martens, 1971: 378; Legg, 1971a: 476; Schaller, 1971: 425; Verner, 1971: figs 43-45, 47; Howes, 1972a: 108; Legg, 1972b: 5; Lagar, 1972a: 21; Muchmore, 1972d: 109; Legg, 1973b: 430, fig. 2j; Ćurčić, 1974a: 25; Legg, 1974a: 324-338, figs 1-10; Jones, 1975a: 88; Klausen, 1975: 64; Legg, 1975a: 66; Legg, 1975d: fig. 2; Frøiland, 1976: 12; Crocker, 1978: 8; Jones, 1978: 94; Judson, 1979a: 63; Schaller, 1979: plate 10.III fig. 7 (as Cheridium [sic] museorum); Thaler, 1979: 52; Gardini, 1980c: 116-117, 122, 129; Jones, 1980c: map 13; Mendel, 1981: 229; Cloudsley-Thompson, 1984: 190; Jedryczkowski, 1985: 79; R.E. Jones, 1985: 69; Lippold, 1985: 40; Schawaller, 1985a: 10; Andersson, Coulianos, Ehnström, Hammarstedt, Imby, Janzon, Lindelöw and Waldén, 1987: 68 (as Cheridium [sic] museorum); Judson, 1987: 369; Kovoor, 1987: 166 (as Cheridium [sic] museorum); El-Hennawy, 1988b: 10; Krumpál and Cyprich, 1988: 42; Legg and Jones, 1988: 91-92, figs 5b, 20a, 20b(a-f); Schaefer, 1988: 99; Chater, 1989: 8; Drogla, 1990: 101; Muchmore, 1990a: 517; Bhattacharyya, 1990a: 245, fig. 1; Harvey, 1991a: 439-440; Harvey, 1991b: 103; Dashdamirov and Schawaller, 1992b: 54; Gärdenfors and Wilander, 1992: 29, fig. 4a; Harvey, 1992c: figs 146-151; P.R. Harvey, 1992: 150; International Commission on Zoological Nomenclature, 1992b: 292; Hogue, 1993: 501 (as Cheiridium muesorum [sic]); Ducháč, 1994: 143; Fowles, 1994: 17; Cooney and Snider, 1995: 241-242; Gardini, 1995a: 6; Jędryczkowski, 1995: 511; Platen, Blick, Bliss, Drogla, Malten, Martens, Sacher and Wunderlich, 1995: 54; Judson, 1997a: 28-29; Legg and O'Connor, 1997: 113, fig. 5; Legg, 1998: 2; Ducháč, 1999a: 14 (as Cheridium [sic] museorum); Alberti, 2000: 312, figs 9a, 19b; Gardini, 2000: 107; Henderickx, 2000c: 67; Petrov, 2000: 63, fig. 1; Šťáhlavský, 2001: 87-88, fig. 8; Delfosse, 2003: 38, unnumbered figure on page 38; Ćurčić, Dimitrijević and Legakis, 2004: 260-261; Drogla, 2004: 6; Drogla and Lippold, 2004: 27; Mahnert, 2004: 464; Ozimec, 2004: 388; Dashdamirov, 2005: 245-247, figs 101-112; Tooren, 2005b: 95; Šťáhlavský, 2006b: 163; Šťáhlavský and Krásný, 2007: 432; Zaragoza, 2007a: 46-47.

Obisium museorum (Leach): Bernard, 1893b: 428; Bernard, 1894: 410.

Not Cheiridium museorum (Leach): Beier, 1955a: 12 (misidentification; see Cheiridium congicum Beier).

 

Type locality: of Chelifer museorum: not stated, presumably England, United Kingdom (possibly Kingsbridge, Devon, England, United Kingdom - see Judson, 1997a: 29).

of Chelifer nepoides: not stated, presumably near Strasbourg, Bas-Rhin, Alsace, France.

Distribution: Algeria; Austria; Belgium; Bulgaria; Croatia; Czech Republic; Democratic Republic of Congo (Zaire); Denmark; Finland; France (mainland); Germany; Greece (mainland); Hungary; India; Ireland; Italy (mainland); Kyrgyzstan; Latvia; Mozambique; Netherlands; Norway; Pakistan; Poland; Romania; Russia (Belgorodskaya Oblast’); Slovakia; South Africa; Spain (mainland); Sweden; Switzerland; Turkey; United Kingdom; U.S.A. (Massachusetts, Michigan); Yugoslavia (Montenegro).

Notes: Franganillo's (1936) record from Cuba is discounted.

Cheiridium nepalense Ćurčić

Cheiridium nepalense Ćurčić, 1980b: 83-84, figs 15-20; Schawaller, 1987c: 212; Harvey, 1991a: 440; Judson, 1997a: 29-30; Beron, 2002: 36.

 

Type locality: Thini Kola, Nepal.

Distribution: Nepal.

Cheiridium nubicum Beier

Cheiridium nubicum Beier, 1962e: 298-300, fig. 2; Cloudsley-Thompson, 1984: 189; El-Hennawy, 1988b: 10; Harvey, 1991a: 440.

 

Type locality: Wadi Halfa, Nil-Oase, Northern, Sudan.

Distribution: Sudan.

Cheiridium perreti Mahnert

Cheiridium perreti Mahnert, 1982b: 123-125, figs 25-29; Harvey, 1991a: 440.

 

Type locality: S. of Mpekatoni, Lamu, Coast, Kenya.

Distribution: Kenya.

Cheiridium reyesi Muchmore

Cheiridium reyesi Muchmore, 1992: 138-139, figs 14-15; Peck, 1998: 23.

 

Type locality: Cot Cave, Kinney County, Texas, U.S.A.

Distribution: U.S.A. (Texas).

Cheiridium saharicum Beier

Cheiridium saharicum Beier, 1965b: 371-373, fig. 5; Harvey, 1991a: 440.

 

Type locality: Mardengai, near Faya (as Largeau), Borkou-Ennedi-Tinesti, Chad.

Distribution: Chad.

Cheiridium simulacrum Chamberlin

Cheiridium (Cheiridium) simulacrum Chamberlin, 1938a: 268-269, figs 2b-d, 2h, 2j-k.

Cheiridium simulacrum Chamberlin: Roewer, 1940: 346; Harvey, 1991a: 440.

 

Type locality: Wahiawa, Oahu Island, Hawaii, U.S.A.

Distribution: U.S.A. (Hawaii).

Cheiridium somalicum Mahnert

Cheiridium somalicum Mahnert, 1984b: 58-60, figs 30-37; Harvey, 1991a: 440.

 

Type locality: Sar Uanle, Jubbada Hoose, Somalia.

Distribution: Somalia.

Cheiridium tumidum Mahnert

Cheiridium tumidum Mahnert, 1982b: 121-123, figs 17-24; Harvey, 1991a: 441.

 

Type locality: SE. of Kogari, Embu, Eastern, Kenya.

Distribution: Kenya.

Genus Cryptocheiridium Chamberlin

Cryptocheiridium Chamberlin, 1931a: 238; Beier, 1932e: 14; Harvey, 1991a: 441.

 

Type species: Cheiridium subtropicum Tullgren, 1907b, by original designation.

Distribution: AFRICA: eastern and southern Africa.

AMERICAS: Caribbean.

ASIA: widespread.

AUSTRALASIA: Australia.

 

Subgenus Cryptocheiridium (Cryptocheiridium) Chamberlin

Cryptocheiridium (Cryptocheiridium) Chamberlin: Harvey, 1991a: 441.

† Cryptocheiridium (Cryptocheiridium) antiquum Schawaller

Cryptocheiridium antiquum Schawaller, 1981d: 3-7, figs 1-12.

Cryptocheiridium (Cryptocheiridium) antiquum Schawaller: Harvey, 1991a: 441.

 

Type locality: Dominican Republic (Miocene Amber).

Distribution: Dominican Republic (Miocene Amber).

Cryptocheiridium (Cryptocheiridium) australicum Beier

Cryptocheiridium australicum Beier, 1969a: 183-185, fig. 10; Richards, 1971: 19, 24, 25, 30, 32, 43; Harvey, 1981b: 244; Harvey, 1985b: 130; Schawaller, 1981d: figs 11-12; Moulds, 2004: 12.

Cryptocheiridium (Cryptocheiridium) australicum Beier: Harvey, 1991a: 441.

 

Type locality: Murra-El-Elevyn Cave [6N-47], Nullarbor Plain, Western Australia, Australia.

Distribution: Australia (Western Australia).

Cryptocheiridium (Cryptocheiridium) elgonense Beier

Cryptocheiridium elgonense Beier, 1955c: 546-548, figs 13-14; Mahnert, 1981c: fig. 2; Mahnert, 1982b: 131-133, figs 34-37; Schawaller, 1981d: figs 11-12; Harvey, 1992c: figs 152-155; Beron, 2000: 162; Beron, 2002: 36.

Cryptocheiridium (Cryptocheiridium) elgonense Beier: Harvey, 1991a: 441.

 

Type locality: Mt Elgon, E. side, Rift Valley, Kenya.

Distribution: Kenya.

Cryptocheiridium (Cryptocheiridium) formosanum (Ellingsen)

Cheiridium formosanum Ellingsen, 1912a: 123-125.

Cryptocheiridium formosanum (Ellingsen): Chamberlin, 1931a: 238; Beier, 1932e: 15; Roewer, 1937: 277; Schawaller, 1981d: figs 11-12.

Cryptocheiridium (Cryptocheiridium) formosanum (Ellingsen): Harvey, 1991a: 441.

 

Type locality: Kao-hsiung (as Takao), Kao-hsiung, Taiwan.

Distribution: Taiwan.

Cryptocheiridium (Cryptocheiridium) kivuense Beier

Cryptocheiridium kivuense Beier, 1959d: 38-39, fig. 18; Beier, 1972a: 9; Mahnert, 1982b: 133; Beron, 2002: 36.

Cryptocheiridium (Cryptocheiridium) kivuense Beier: Harvey, 1991a: 441.

 

Type locality: Itombwe, Sud-Kivu, Democratic Republic of Congo (Zaire).

Distribution: Democratic Republic of Congo (Zaire); Kenya.

Cryptocheiridium (Cryptocheiridium) lucifugum Beier

Cryptocheiridium lucifugum Beier, 1963c: 51-52, fig. 1; McClure, Lim and Winn, 1967: 425; Tenorio and Muchmore, 1982: 378.

Cryptocheiridium (Cryptocheiridium) lucifugum Beier: Harvey, 1991a: 441.

 

Type locality: Cavern A, Batu Caves, near Kuala Lumpur, Selangor, Malaysia.

Distribution: Malaysia (mainland).

Cryptocheiridium (Cryptocheiridium) philippinum Beier

Cryptocheiridium philippinum Beier, 1977a: 187-188, fig. 1; Schawaller, 1981d: figs 11-12; Strinati, 1983: 30.

Cryptocheiridium (Cryptocheiridium) philippinum Beier: Harvey, 1991a: 441-442.

 

Type locality: Cueva Basilen, Pagbilao, Quezon, Philippines.

Distribution: Philippines.

Cryptocheiridium (Cryptocheiridium) salomonense Beier

Cryptocheiridium salomonense Beier, 1970g: 319-320, fig. 2; Beier, 1971b: 367.

Cryptocheiridium (Cryptocheiridium) salomonense Beier: Harvey, 1991a: 442; Judson, 1997a: 37.

 

Type locality: N. of Kuzi, Kolombangara Island, Western, Solomon Islands.

Distribution: Papua New Guinea; Solomon Islands.

Cryptocheiridium (Cryptocheiridium) somalicum Callaini

Cryptocheiridium somalicum Callaini, 1985: 182-186, figs 1, 2a-g; Messana, Chelazzi and Baccetti, 1985: 334.

Cryptocheiridium (Cryptocheiridium) somalicum Callaini: Harvey, 1991a: 442.

 

Type locality: Mugdile Cave, near Baardheere, Gedo, Somalia.

Distribution: Somalia.

Cryptocheiridium (Cryptocheiridium) subtropicum (Tullgren)

Cheiridium subtropicum Tullgren, 1907b: 218-220, figs 5a-b, 5d-f (as Chiridium [sic] subtropicum); Ellingsen, 1912b: 86, 104; Godfrey, 1923: 98-99; Godfrey, 1927: 18.

Cryptocheiridium subtropicum (Tullgren): Chamberlin, 1931a: 238; Beier, 1932e: 14-15, fig. 10; Vachon, 1934i: 414-415; Roewer, 1937: 277; Beier, 1958a: 171; Beier, 1964k: 69; Schawaller, 1981d: figs 11-12; Haddad, Dippenaar-Schoeman and Wesołowska, 2006: 21.

Cryptocheiridium (Cryptocheiridium) subtropicum (Tullgren): Harvey, 1991a: 442.

 

Type locality: Mfolozi (as Umfolozi), KwaZulu-Natal Province, South Africa.

Distribution: South Africa.

Subgenus Cryptocheiridium (Cubanocheiridium) Dumitresco and Orghidan

Cryptocheiridium (Cubanocheiridium) Dumitresco and Orghidan, 1981: 77; Harvey, 1991a: 442.

 

Type species: Cryptocheiridium (Cubanocheiridium) elegans Dumitresco and Orghidan, 1981, by original designation.

Cryptocheiridium (Cubanocheiridium) elegans Dumitresco and Orghidan

Cryptocheiridium (Cubanocheiridium) elegans Dumitresco and Orghidan, 1981: 77-84, figs 1a-b, 2a-b, 3a-f, 4a, 5a, 6a, 7a-f; Armas and Alayón, 1984: 11; Harvey, 1991a: 442.

 

Type locality: Cueva La Pluma, Corral Nuevo, Matanzas Municipality, Matanzas, Cuba.

Distribution: Cuba.

† Genus Electrobisium Cockerell

Electrobisium Cockerell, 1917: 360; Harvey, 1991a: 334; Judson, 2000: 80-81.

 

Type species: Electrobisium acutum Cockerell, 1917, by original designation.

† Electrobisium acutum Cockerell

Electrobisium acutum Cockerell, 1917: 360, fig. 1; Schawaller, 1978: 3; Morris, 1980: 36; Harvey, 1991a: 334; Judson, 1997a: 7; Judson, 2000: 80-81, figs 1-4.

 

Type locality: Hukawng (as Hukong) Valley, Kachin, Myanmar (amber fossil, possibly Upper Cretaceous).

Distribution: Myanmar (amber fossil; possibly Upper Cretaceous).

Genus Neocheiridium Beier

Neocheiridium Beier, 1932e: 9; Harvey, 1991a: 442.

Neocheiridium (Austrocheiridium) Vitali-di Castri, 1962: 134 (synonymised by Mahnert, 1982b: 130).

 

Type species: of Neocheiridium: Cheiridium corticum Balzan, 1887b, by original designation.

of Neocheiridium (Austrocheiridium): Neocheiridium (Austrocheiridium) chilense Vitali-di Castri, 1962, by original designation.

Distribution: AFRICA: eastern Africa.

AMERICAS: widespread.

Neocheiridium africanum Mahnert

Neochiridium sp.: Mahnert, 1981c: fig. 2.

Neocheiridium africanum Mahnert, 1982b: 127-128, figs 38-45; Harvey, 1991a: 442.

 

Type locality: Warges, Rift Valley, Kenya.

Distribution: Kenya.

Neocheiridium beieri Vitali-di Castri

Neocheiridium beieri Vitali-di Castri, 1962: 131-133, figs 5a-b; Beier, 1964b: 341; Vitali-di Castri, 1966a: fig. 3; Vitali-di Castri, 1966b: figs 6, 9; Cekalovic, 1984: 19; Harvey, 1991a: 442-443.

Neocheiridium (Neocheiridium) beieri Vitali-di Castri: Vitali-di Castri, 1969b: 266.

 

Type locality: Hacienda La Rinconada de Maipú, Quebrada La Plata, Región Metropolitana, Chile.

Distribution: Chile.

Neocheiridium chilense Vitali-di Castri

Neocheiridium (Austrocheiridium) chilense Vitali-di Castri, 1962: 134-138, figs 6a-d; Beier, 1964b: 341; Vitali-di Castri, 1966a: fig. 3; Vitali-di Castri, 1966b: figs 5, 9; Vitali-di Castri, 1969b: 266-267; Cekalovic, 1984: 19.

Neocheiridium chilense Vitali-di Castri: Harvey, 1991a: 443.

 

Type locality: Hacienda La Rinconada de Maipú, Quebrada La Plata, Región Metropolitana, Chile.

Distribution: Chile.

Neocheiridium corticum (Balzan)

Cheiridium corticum Balzan, 1887b: no pagination, figs; Balzan, 1890: 410-411, figs 1, 1a-c; Balzan, 1892: 549.

Apocheiridium (?) corticum (Balzan): Chamberlin, 1931a: 238.

Neocheiridium corticum (Balzan): Beier, 1932e: 9-10, fig. 6; Vachon, 1934i: 414; Roewer, 1937: 277; Mello-Leitão, 1939a: 117; Mello-Leitão, 1939b: 615; Beier, 1977c: 103; Mahnert and Aguiar, 1986: 499-503, figs 1-8; Harvey, 1991a: 443; Aguiar and Bührnheim, 1992b: 429; Baert, Maelfait and Desender, 1995: 18; Aguiar, Arruda and Bührnheim, 1997: 83; Aguiar and Bührnheim, 1998a: 383-384; Aguiar and Bührnheim, 1998b: 454; Poinar, Ćurčić and Cokendolpher, 1998: 85; Mahnert and Adis, 2002: 378.

 

Type locality: Rio Apa, Paraguay.

Distribution: Argentina; Brazil (Roraima); Ecuador (Galapagos Islands); Paraguay.

Neocheiridium galapagoense Beier

Neocheiridium galapagoense Beier, 1978c: 540-541, fig. 5; Harvey, 1991a: 443; Baert, Maelfait and Desender, 1995: 18.

 

Type locality: Isla Pinzón, Galapagos Islands, Galápagos, Ecuador.

Distribution: Ecuador (Galapagos Islands).

Neocheiridium pusillum Mahnert

Neocheiridium pusillum Mahnert, 1982b: 128-130, figs 46-55; Mahnert, Adis and Bührnheim, 1987: fig. 11b; Harvey, 1991a: 443.

 

Type locality: 10 km S. of Garsen, Tana River, Coast, Kenya.

Distribution: Kenya.

Neocheiridium strinatii Mahnert and Aguiar

Neocheiridium strinatii Mahnert and Aguiar, 1986: 503-505, figs 9-12; Harvey, 1991a: 443; Muchmore, 1993a: 92; Aellen and Strinati, 1997: 276.

 

Type locality: Grot van San Pedro, Curaçao, Netherlands Antilles.

Distribution: Netherlands Antilles (Aruba; Curaçao).

Neocheiridium tenuisetosum Beier

Neocheiridium tenuisetosum Beier, 1959e: 202, fig. 14; Harvey, 1991a: 443.

 

Type locality: Lago Trebol, San Carlos de Bariloche, Río Negro, Argentina.

Distribution: Argentina.

Neocheiridium triangulare Mahnert and Aguiar

Neocheiridium triangulare Mahnert and Aguiar, 1986: 505-508, figs 13-17; Harvey, 1991a: 443; Aguiar and Bührnheim, 1994: 74; Aguiar and Bührnheim, 1998b: 454; Poinar, Ćurčić and Cokendolpher, 1998: 89; Mahnert and Adis, 2002: 378.

 

Type locality: Rio Urubu, Amazonas, Brazil.

Distribution: Brazil (Acre, Amazonas, Pará, Rondonia).

Genus Nesocheiridium Beier

Nesocheiridium Beier, 1957d: 27; Harvey, 1991a: 443.

 

Type species: Nesocheiridium stellatum Beier, 1957d, by original designation.

Distribution: ASIA: Micronesia.

Nesocheiridium stellatum Beier

Nesocheiridium stellatum Beier, 1957d: 28, figs 13a-c; Harvey, 1991a: 444.

 

Type locality: Mt Magpi, Pidos Kalahe, Saipan, Northern Mariana Islands.

Distribution: Northern Mariana Islands.

Subfamily PYCNOCHEIRIDIINAE Beier

Pycnocheiridiinae Beier, 1964k: 66.

Genus Pycnocheiridium Beier

Pycnocheiridium Beier, 1964k: 65-66; Harvey, 1991a: 444.

 

Type species: Pycnocheiridium mirum Beier, 1964k, by original designation.

Distribution: AFRICA: South Africa.

Pycnocheiridium mirum Beier

Pycnocheiridium mirum Beier, 1964k: 66-67, fig. 25; Harvey, 1991a: 444.

 

Type locality: Wilton Farm, near Alicedale, Eastern Cape, South Africa.

Distribution: South Africa.

CATALOGUE

 

Family Cheiridiidae Hansen 1894