Pseudoscorpions of the World

Last updated: 19 January 2009

Version 1.2, 2009

Family SYARINIDAE Chamberlin

Syarinidae Chamberlin, 1930: 38; Chamberlin, 1931a: 218-219; Beier, 1932a: 162-163; Beier, 1932g: 183; Roewer, 1937: 253; Hoff, 1956b: 9; Hoff, 1964a: 6; Muchmore, 1979a: 195; Muchmore, 1982a: 97; Harvey, 1991a: 417; Harvey, 1991d: 495; Gärdenfors and Wilander, 1992: 29; Harvey, 1992c: 1412-1414; Harvey, 1996a: 258.

Syarininae Chamberlin, 1930: 39; Chamberlin, 1931a: 219; Beier, 1932a: 163; Chamberlin, 1938b: 109-110; Hoff, 1956b: 10.

Subfamily CHITRELLINAE Beier

Chitrinae Chamberlin, 1930: 40; Chamberlin, 1931a: 219.

Chitrellinae Beier, 1932a: 165; Chamberlin, 1938b: 110; Hoff, 1956b: 20; Hoff, 1964a: 7.

Microcreagrellinae Beier, 1963b: 222.

Genus Aglaochitra Chamberlin

Aglaochitra Chamberlin, 1952: 268-269; Harvey, 1991a: 417.

 

Type species: Aglaochitra rex Chamberlin, 1952, by original designation.

Distribution: NORTH AMERICA: western U.S.A.

Aglaochitra rex Chamberlin

Aglaochitra rex Chamberlin, 1952: 269-274, figs 2a-f, 3a-h; Hoff, 1958: 14; Muchmore, 1971a: 90; Muchmore, 1990a: 513; Harvey, 1991a: 417; Francke and Villegas-Guzmán, 2006: 294.

 

Type locality: Red Hill, Frances Simes Hastings Natural History Reservation, 39 miles S. of Monterey, Monterey County, California, U.S.A.

Distribution: U.S.A. (California).

Genus Chitrella Beier

Chitra Chamberlin, 1930: 40 (preoccupied by Chitra Gray, 1844).

Chitrella Beier, 1932a: 165; Hoff, 1956b: 20-21; Muchmore, 1973e: 183; Harvey, 1991a: 418; Muchmore, 1992: 134 (replacement name for Chitra Chamberlin, 1930).

 

Type species: Chitra cala Chamberlin, 1930, by original designation.

Distribution: NORTH AMERICA: U.S.A.

Chitrella archeri Malcolm and Chamberlin

Chitrella archeri Malcolm and Chamberlin, 1960: 3-7, figs 1a-e; Barr, 1961: 34; Harvey, 1991a: 418.

 

Type locality: Wonder Cave, Monteagle, Marion County, Tennessee, U.S.A.

Distribution: U.S.A. (Tennessee).

Chitrella cala (Chamberlin)

Chitra cala Chamberlin, 1930: 41, figs 2x, 2jj; Chamberlin, 1931a: 82, figs 13k, 17o, 33p, 36c, 40r, 43l-m, 47z, 57.

Chitrella cala (Chamberlin): Beier, 1932a: 165-166; Beier, 1932g: fig. 254; Roewer, 1936: fig. 30g; Roewer, 1937: 254; Hoff, 1958: 14; Hoff, 1959b: 4, 26, 34, 60; Malcolm and Chamberlin, 1960: 11-19, figs 4, 5a-e; Harvey, 1991a: 418; Harvey and Volschenk, 2007b: 369, figs 1-4.

 

Type locality: San Francisquito Creek, Stanford University, Palo Alto, Santa Clara County, California, U.S.A.

Distribution: U.S.A. (California, Utah).

Chitrella cavicola (Packard)

Obisium cavicola Packard, 1884: 202-203, fig. 1 (as Obisium cavirola [sic]; Packard, 1888: 42, fig. 11; Banks, 1895a: 11; Coolidge, 1908: 113 (as Obisium cavicala [sic]).

Microcreagris (?) cavicola (Packard): Beier, 1932a: 157; Roewer, 1937: 252; Hoff, 1958: 12.

Microcreagris cavicola (Packard): Wolf, 1938: 619.

Chitrella cavicola (Packard): Muchmore, 1962: 5; Holsinger, 1963: 34; Muchmore, 1963a: 11-14, figs 7-8; Holsinger, 1964: 63; Muchmore, 1973e: 184-187, figs 1-4; Holsinger and Culver, 1988: 43; Muchmore, 1990a: fig. 18.38; Harvey, 1991a: 418

 

Type locality: New Market Cave, Shenandoah County, Virginia, U.S.A.

Distribution: U.S.A. (Virginia, West Virginia).

Chitrella elliotti Muchmore

Chitrella elliotti Muchmore, 1992: 137, figs 12-13

 

Type locality: Felton Cave, Sutton County, Texas, U.S.A.

Distribution: U.S.A. (Texas).

Chitrella major Muchmore

Chitrella major Muchmore, 1992: 135-137, fig. 11.

 

Type locality: Fern Cave, Val Verde County, Texas, U.S.A.

Distribution: U.S.A. (Texas).

Chitrella muesebecki Malcolm and Chamberlin

Chitrella muesebecki Malcolm and Chamberlin, 1960: 9-11, figs 3a-d; Harvey, 1991a: 418.

 

Type locality: Roane County, Tennessee, U.S.A.

Distribution: U.S.A. (Tennessee).

Chitrella regina Malcolm and Chamberlin

Chitrella regina Malcolm and Chamberlin, 1960: 7-9, figs 2a-d; Muchmore, 1973e: 187-189, fig. 5; Harvey, 1991a: 418-419; Drewry, 1989: 566.

 

Type locality: Coffman's Cave, Frankford (as Frankfurt), Greenbrier County, West Virginia, U.S.A.

Distribution: U.S.A. (West Virginia).

Chitrella superba Muchmore

Chitrella superba Muchmore, 1973e: 189-190, figs 6-8; Holsinger and Culver, 1988: 43, fig. 15; Harvey, 1991a: 419.

 

Type locality: Madden's Cave, Shenandoah County, Virginia, U.S.A.

Distribution: U.S.A. (Virginia).

Chitrella transversa (Banks)

Obisium transversum Banks, 1909b: 307.

Microcreagris transversa (Banks): Beier, 1932a: 157; Roewer, 1937: 252.

Chitrella transversa (Banks): Hoff, 1956b: 21-24, figs 11-12; Hoff, 1958: 14; Hoff, 1959b: 4, etc.; Hoff, 1961: 434-435; Malcolm and Chamberlin, 1960: 19; Muchmore, 1973e: 190-191, figs 9-10; Harvey, 1991a: 419.

 

Type locality: Pecos, San Miguel County, New Mexico, U.S.A.

Distribution: U.S.A. (Colorado, New Mexico).

Chitrella welbourni Muchmore

Chitrella sp.: Welbourn, 1978: 31.

Chitrella welbourni Muchmore, 1992: 134-135, figs 8-10.

 

Type locality: Ogle Cave (Ogle Section), Carlsbad Caverns National Park, Eddy County, New Mexico, U.S.A.

Distribution: U.S.A. (New Mexico).

Genus Chitrellina Muchmore

Chitrellina Muchmore, 1996g: 145.

 

Type species: Chitrellina chiricahuae Muchmore, 1996g, by original designation.

Distribution: NORTH AMERICA: U.S.A.

Chitrellina chiricahuae Muchmore

Chitrellina chiricahuae Muchmore, 1996g: 147-148, figs 1-7; Reddell, 2005: 560.

 

Type locality: Spinks (or Sphinx) Cave, SW. of San Simon, Chiricahua Mountains, Cochise County, Arizona, U.S.A.

Distribution: U.S.A. (Arizona).

Genus Hadoblothrus Beier

Hadoblothrus Beier, 1952d: 106; Beier, 1963b: 226-227; Harvey, 1991a: 419.

 

Type species: Parablothrus gigas Caporiacco, 1951f, by original designation.

Distribution: EUROPE: Greece; Italy.

Hadoblothrus aegeus Beron

Hadoblothrus aegeus Beron, 1985: 67, figs 3-7; Harvey, 1991a: 419.

 

Type locality: Zoodochos I and II caves, Santorin, Kyklades, Greece; and Agios Ioannis Cave, Iraklia, Kyklades, Greece.

Distribution: Greece (Kyklades).

Hadoblothrus gigas (Caporiacco)

Parablothrus gigas Caporiacco: Ruffo, 1950: 60 (not seen) (nomen nudum).

Parablothrus gigas Caporiacco, 1951f: 97, fig. 7.

Hadoblothrus gigas (Caporiacco): Beier, 1952d: 106-107, fig. 2; Baggini, 1961: 24; Beier, 1963b: 226, fig. 233; Lazzeroni, 1969a: 333; Beier, 1970f: 322, fig. 2; Lazzeroni, 1970a: 207; Gardini, 1980c: 116, 130; Mahnert, 1980d: 35-37, figs 17-23; Muchmore, 1982c: 218; Inzaghi, 1983: 47-48; Harvey, 1991a: 419; Gardini, 1995a: 5; Gardini, 2000: 103-104.

 

Type locality: L'Abisso di Castro Marino (as "Grotta l'Abisso, Castromarina"), Castro, Puglia, Italy.

Distribution: Italy (mainland).

Genus Microcreagrella Beier

Microcreagrella Beier, 1961a: 70-72; Beier, 1963b: 224; Harvey, 1991a: 426.

 

Type species: Obisium caecum Simon, 1883, by original designation.

Distribution: EUROPE: Azores; Madeira; Islas Canarias.

Microcreagrella caeca (Simon)

Obisium caecum Simon, 1883: 279-280.

Ideobisium (Ideoblothrus) caecum (Simon): Balzan, 1892: 542.

Microcreagris caeca (Simon): Beier, 1932a: 157; Roewer, 1937: 252; Vachon, 1940b: 108-110, figs 1-6; Vachon, 1961: 102, 103 (as Microcreagris coeca [sic]).

Microcreagrella caeca (Simon): Beier, 1961a: 72-74, fig. 2; Beier, 1970f: 322, fig. 2; Beier, 1976a: 24; Pieper, 1981: 2-3; Harvey, 1991a: 426; Pieper, 1991: 218.

 

Type locality: Ponta Delgada (as Ponta-Delgada), São Miguel, Azores, Portugal.

Distribution: Portugal (Azores, Madeira); Spain (Islas Canarias).

Microcreagrella caeca caeca (Simon)

Microcreagrella caeca caeca (Simon): Beier, 1963b: 225; Muchmore, 1982c: 218; Mahnert, 1997: 567.

 

Distribution: Portugal (Azores, Madeira); Spain (Islas Canarias).

Microcreagrella caeca madeirensis Beier

Microcreagrella caeca madeirensis Beier, 1963b: 225, fig. 231; Beier, 1970f: 322, fig. 2; Beier, 1976a: 24; Harvey, 1991a: 426; Mahnert, 1997: 567.

 

Type locality: Madeira, Madeira, Portugal.

Distribution: Portugal (Madeira).

Genus Microcreagrina Beier

Microcreagrina Beier, 1961b: 29-30; Beier, 1963b: 223; Harvey, 1991a: 427.

 

Type species: Microcreagris maroccana Beier, 1931c (junior synonym of Ideobisium hispanicum Ellingsen, 1910a), by original designation.

Distribution: AFRICA: north Africa; Middle East.

EUROPE: Islas Canarias.

Microcreagrina cavicola Mahnert

Microcreagrina cavicola Mahnert, 1993b: 983-985, figs 31-33.

 

Type locality: Cueva Tacande, Tacande de Arriba, Insel La Palma, Islas Canarias, Spain.

Distribution: Spain (Islas Canarias).

Microcreagrina hispanica (Ellingsen)

Ideobisium hispanicum Ellingsen, 1910a: 394; Navás, 1918: 112; Navás, 1925: 114.

Ideobisium (Ideoblothrus) hispanicum Ellingsen: Nonidez, 1917: 32.

Ideoroncus cambridgei (L. Koch): Beier, 1930f: 71 (misidentification).

Microcreagris maroccana Beier, 1931c: 301-302; Beier, 1932a: 155, fig. 187; Roewer, 1937: 251; Vachon, 1940g: 144 (synonymised by Beier, 1970a: 45).

Microcreagris hispanica (Ellingsen): Beier, 1932a: 155-156; Roewer, 1937: 251; Beier, 1939f: 189; Beier, 1963b: 212.

Microcreagris parisi Vachon, 1937e: 107-109, figs 1-2; Roewer, 1940: 344 (synonymised by Mahnert, 1976d: 206).

Microcreagrina maroccana (Beier): Beier, 1961b: 30; Beier, 1963b: 223-224, fig. 230; Beier, 1965c: 377; Lazzeroni, 1969c: 237-238.

Microcreagrina hispanica (Ellingsen): Beier, 1970a: 45; Beier, 1970f: 322, fig. 2; Beier, 1975a: 56; Beier, 1976a: 24; Orghidan, Dumitresco and Georgesco, 1975: 30; Mahnert, 1976c: 206; Mahnert, 1977d: 97; Mahnert, 1980a: 259; Callaini, 1982a: 17, 20, fig. 2c; Mahnert, 1982f: 297; Muchmore, 1982c: 218; Callaini, 1983c: 292; Bacallado, 1984: 110; Callaini, 1988b: 46; Mahnert, 1989b: 42; Harvey, 1991a: 427; Pieper, 1991: 218, 219; Mahnert, 1993a: 357; Gardini, 1995a: 5; Mahnert, 1997: 566-567; Gama, Sousa, Ferreira and Barrocas, 2000: 91; Gardini, 2000: 103; Zaragoza, 2007a: 46.

Microcreagrina cf. hispanica (Ellingsen): Medina and Oromi, 1990: 89.

Microcreagrina gomerae Mahnert, 1993b: 980-981, figs 21-25 (synonymised by Mahnert, 1997: 566).

 

Type locality: of Ideobisium hispanicum: Algeciras, Cádiz, Andalucía, Spain.

of Microcreagris maroccana: Korifla, Rabat-Salé, Morocco.

of Microcreagris parisi: Skikda (as Philippeville), Constantine, Algeria.

Distribution: Algeria; Italy (mainland, Sardinia, Sicily); Lebanon; Malta; Morocco; Portugal (mainland, Madeira); Spain (mainland, Islas Baleares, Islas Canarias). of Microcreagrina gomerae: El Cedro, Gomera, Islas Canarias, Spain.

Microcreagrina madeirensis Mahnert

Microcreagrina madeirensis Mahnert, 1993b: 986-988, figs 40-42.

 

Type locality: S. of Pedra Branchas, Ponta de S. Lourenço, Madeira, Portugal.

Distribution: Portugal (Madeira).

Microcreagrina subterranea Mahnert

Microcreagrina subterranea Mahnert, 1993b: 981, figs 26-30; Mahnert, 1997: 567; Mahnert, 2002: 778.

 

Type locality: Cueva Felipe Reventón, 3 km S. of Icod, Tenerife, Islas Canarias, Spain.

Distribution: Spain (Islas Canarias).

Genus Pararoncus Chamberlin

Pararoncus Chamberlin, 1938a: 260; Roewer, 1940: 344; Ćurčić, 1979a: 176-177; Harvey, 1991a: 393.

 

Type species: Pararoncus histrionicus Chamberlin, 1938a, by original designation.

Distribution: ASIA: Japan; Korea.

Pararoncus chamberlini (Morikawa)

Roncus (Roncus) troglophilus Morikawa, 1957a: 360, figs 2c, 2g (junior primary homonym of Roncus (Parablothrus) troglophilus Beier, 1931a).

Roncus (Roncus) chamberlini Morikawa, 1960: 117-118; Morikawa, 1962: 418, fig. 4; Sato, 1994b: 80 (replacement name for Roncus (Roncus) troglophilus Morikawa).

Roncus (Roncus) chamberlini chamberlini Morikawa: Morikawa, 1960: 118, plate 6 figs 2-3, plate 9 figs 7, 9; Morikawa, 1962: fig. 4; Sato, 1994b: 80.

Pararoncus troglophilus (Morikawa): Ćurčić, 1979a: 174.

Pararoncus chamberlini (Morikawa): Harvey, 1991a: 393.

 

Type locality: Goyomatsu-dani-dô Cave, Dorogawa, Tenkawa-mura, Nara Prefecture, Japan.

Distribution: Japan.

Pararoncus histrionicus Chamberlin

Pararoncus histrionicus Chamberlin, 1938a: 261-263, figs 1a-g; Roewer, 1940: 344, fig. 260; Ćurčić, 1979a: 169; Harvey, 1991a: 393.

Roncus (Roncus) japonicus (Ellingsen): Morikawa, 1960: 117-118 (misidentification, in part).

 

Type locality: Japan.

Distribution: Japan.

Pararoncus japonicus (Ellingsen)

Obisium (Roncus) japonicum Ellingsen, 1907c: 10-12.

Roncus japonicus (Ellingsen): Kishida, 1929a: 39; Sato, 1978c: 40; Sato, 1979d: 43; Sato, 1982b: 58, figs 1-3; Sato, 1982c: 32; Sato, 1983b: 16, fig. 2; Sato, 1984d: fig. 2; Sato, 1986: 95; Sato, 1988: 12, figs 1, 3; Sato, Takano and Nakajima, 1988: 23; Sakayori, 1990: 33, fig. 2C; Nakajima, Takano, Sato and Tashiro, 1991: 16; Sato, 1991a: 23, fig. 68-3; Hong and Kim, 1996: 134-136, figs 1a-d, plates a-h; Sato and Yamamoto, 2000: 666.

Roncus (Roncus) (?) japonicus (Ellingsen): Beier, 1932a: 129 (as Roncus (Roncus) (?) japonicum [sic]); Roewer, 1937: 249 (as Roncus (Roncus) (?) japonicum [sic]).

Roncus (Roncus) japonicus (Ellingsen): Morikawa, 1952b: 253-254, figs 7-8; Morikawa, 1955a: 222; Morikawa, 1960: 117-118, plate 2 fig. 4, plate 7 fig. 11 (in part; see Pararoncus histrionicus Chamberlin); Morikawa, 1962: 418, figs 2, 4; Sato, 1979a: 94-95, figs 7a-k, 8a-e, plates 3, 12; Sato, 1994b: 80.

Pararoncus japonicus (Ellingsen): Ćurčić, 1979a: 172-174; Sato, 1980d: plate 2; Harvey, 1991a: 393; Nunomura, Hirauchi and Sato, 1997: 13; Sakayori, 1998a: 300.

 

Type locality: Yamanaka, Suruga-wan (as Suruga), Yamanashi Prefecture, Japan.

Distribution: Japan; South Korea.

Pararoncus oinuanensis (Morikawa)

Roncus (Roncus) troglophilus oinuanensis Morikawa, 1957a: 360, figs 2d, 2h.

Roncus (Roncus) chamberlini oinuanensis Morikawa: Morikawa, 1960: 118; Sato, 1994b: 80.

Pararoncus oinuanensis (Morikawa): Ćurčić, 1979a: 174-176; Harvey, 1991a: 393.

 

Type locality: Oinu-ana Cave, Nakakoshizawa, Uéno-mura, Gumma Prefecture, Japan.

Distribution: Japan.

Pararoncus rakanensis (Morikawa)

Roncus (Roncus) uénoi rakanensis Morikawa, 1957a: 361-363, figs 3b-c, 3e.

Roncus (Parablothrus) rakanensis Morikawa: Morikawa, 1960: 119, plate 2 fig. 5, plate 6 figs 5-6, plate 9 fig. 5; Sato, 1994b: 80.

Pararoncus rakanensis (Morikawa): Ćurčić, 1979a: 170-172, figs 1-7; Harvey, 1991a: 394.

 

Type locality: Rakan-ana Cave, Uchiko-chô, Ehime Prefecture, Japan.

Distribution: Japan.

Pararoncus uenoi (Morikawa)

Roncus (Roncus) uenoi Morikawa, 1957a: 361, figs 3a, 3d (as Roncus (Roncus) uénoi).

Roncus (Parablothrus) uenoi Morikawa: Morikawa, 1960: 119, plate 6 fig. 9, plate 9 fig. 6; Sato, 1994b: 80.

Pararoncus uenoi (Morikawa): Ćurčić, 1979a: 169-170; Harvey, 1991a: 394.

 

Type locality: Yôzawa-dô Cave, Komiya-mura, Tokyo Prefecture, Japan.

Distribution: Japan.

Pararoncus yosii (Morikawa)

Roncus (Roncus) chamberlini yosii Morikawa, 1960: 118; Sato, 1994b: 80.

Pararoncus yosii (Morikawa): Ćurčić, 1979a: 176; Harvey, 1991a: 394.

 

Type locality: Shirataki-do, Iwato-mura, Miyazaki Prefecture, Japan.

Distribution: Japan.

Genus Pseudoblothrus Beier

Pseudoblothrus Beier, 1931a: 21; Beier, 1932a: 135; Beier, 1963b: 227-228; Harvey, 1991a: 428.

 

Type species: Ideoblothrus roszkovskii Redikorzev, 1918, by original designation.

Distribution: EUROPE: widespread; including Ukraine.

Pseudoblothrus ellingseni (Beier)

Obisium (Blothrus) torrei Simon: Ellingsen, 1905d: 9-11 (misidentification).

Blothrus peyerimhoffi Simon, 1905: 282-283 (misidentification, in part).

Obisium (Blothrus) ellingseni Beier, 1929a: 363.

Neobisium (Blothrus) ellingseni (Beier): Beier, 1932a: 113; Roewer, 1937: 248.

Neobisium ellingseni (Beier): Wolf, 1938: 611.

Pseudoblothrus ellingseni (Beier): Beier, 1963b: 228-229; Gardini, 1980c: 116, 119; Mahnert, 1980d: 30-32, figs 13-14; Muchmore, 1982c: 218; Bologna and Taglianti, 1985: 228-229, fig. 9; Harvey, 1991a: 428; Gardini, 1995a: 5; Gardini, 1996: 158; Gardini, 2000: 104.

Pseudoblothrus sp. prope peyerimhoffi (Simon): Vigna Taglianti, 1969: 267 (misidentification).

Pseudoblothrus ellingseni (Beier) spp.: Mahnert, 1980d: 32-33.

 

Type locality: Grotta di Bossea, Frabosa Soprano, Piemonte e Val d'Aosta, Italy.

Distribution: Italy (mainland).

Pseudoblothrus ljovuschkini Krumpál

Pseudoblothrus ljovuschkini Krumpál, 1984a: 642-645, figs 11-16; Harvey, 1991a: 428.

 

Type locality: Peshtshera Egiz, Karabijajra, Krym, Ukraine.

Distribution: Ukraine.

Pseudoblothrus oromii Mahnert

Pseudoblothrus oromii Mahnert, 1990a: 168-169, figs 1-10; Oromi, Martin, Ashmole and Ashmole, 1990: 99.

 

Type locality: Gruta da Beira, Sao Jorge, Azores, Portugal.

Distribution: Portugal (Azores).

Pseudoblothrus peyerimhoffi (Simon)

Blothrus peyerimhoffi Simon, 1905: 282-283 (in part; see Pseudoblothrus ellingseni (Beier)); Peyerimhoff, 1906: 206, 220; Vachon, 1938a: figs 37i, 38a.

Obisium (Blothrus) peyerimhoffi (Simon): Jeannel, 1926: 154.

Neobisium (Blothrus) peyerimhoffi (Simon): Beier, 1932a: 113; Roewer, 1937: 248.

Neobisium peyerimhoffi (Simon): Wolf, 1938: 613.

Pseudoblothrus peyerimhoffi (Simon): Vachon, 1945a: 230-232, figs 1-7; Vachon, 1952c: 536-537; Balazuc, 1962: 106; Beier, 1963b: 229; Beier, 1970f: 322, fig. 2; Mahnert, 1980d: 33-34, figs 15-16; Leclerc, 1983b: 25; Leclerc, 1984a: 56; Heurtault, 1986c: 28; Harvey, 1991a: 428; Heurtault, 1994b: fig. 2; Pascutto, 1998: 46 (not seen); Gardini, 2000: 104; Delfosse, 2003: 37.

Not Pseudoblothrus sp. prope peyerimhoffi (Simon): Vigna Taglianti, 1969: 267 (misidentification; see Pseudoblothrus ellingseni (Beier)).

 

Type locality: Grotte de Mélan, Méailles, Alpes-de-Haute-Provence, Provence-Alpes-Côtes-d'Azur, France.

Distribution: France (mainland); Italy (mainland).

Pseudoblothrus regalini Inzaghi

Pseudoblothrus regalini Inzaghi, 1983: 38-46, figs 1-12; Harvey, 1991a: 428; Gardini, 1995a: 5; Gardini, 2000: 105.

 

Type locality: NE. of Monte di Grone, Bergamo, Lombardia, Italy.

Distribution: Italy (mainland).

Pseudoblothrus roszkovskii (Redikorzev)

Ideoblothrus roszkovskii Redikorzev, 1918: 94-96, figs 3, 4, 4a; Pliginsky, 1927: 172.

Pseudoblothrus roszkovskii (Redikorzev): Beier, 1931a: 21-22; Beier, 1932a: 136, fig. 167; Roewer, 1937: 249; Wolf, 1938: 618; Roewer, 1940: 345; Beier, 1963b: 228, fig. 234; Beier, 1970f: 322, fig. 2; Muchmore, 1982c: 218; Harvey, 1991a: 428.

 

Type locality: Suuk-Koba Cave, Krym, Ukraine.

Distribution: Ukraine.

Pseudoblothrus strinatii Vachon

Pseudoblothrus strinatii Vachon, 1954b: 212-217, figs 1-14; Aellen and Strinati, 1956: 192; Strinati, 1957: 62; Aellen and Strinati, 1962: 46; Beier, 1963b: 229; Strinati, 1966: 70-72, 326, fig.; Beier, 1970f: 322, fig. 2; Aellen, 1976: 23; Vachon, 1976: 243, 244; Bourne and Cherix, 1981: 24, fig. 1; Aellen, 1986: 33; Harvey, 1991a: 429.

 

Type locality: Grotte de Pertuis, Jura Neuchâtelois, Neuchâtel, Switzerland.

Distribution: Switzerland.

Pseudoblothrus thiebaudi Vachon

Pseudoblothrus thiebaudi Vachon, 1969: 387-390, figs 1-10; Strinati, 1966: 72, 327, fig.; Vachon, 1976: 244, 245; Thaler, 1980: 391; Harvey, 1991a: 429.

 

Type locality: Neuenburgerhöhle (as Neuenbürgenhöhle), LU2, Luzern, Switzerland.

Distribution: Switzerland.

Pseudoblothrus vulcanus Mahnert

Pseudoblothrus vulcanus Mahnert, 1990a: 169-170, figs 11-13; Oromi, Martin, Ashmole and Ashmole, 1990: 99; Mahnert, 2002: 779.

 

Type locality: Gruta das Agulhas, Terceira, Azores, Portugal.

Distribution: Portugal (Azores).

Subfamily IDEOBISIINAE Banks

Ideobisiini Banks, 1895a: 2 (as Ideobisini [sic]).

Ideobisiinae Banks: Comstock, 1913: 50 (as Ideobsiidae [sic]); Moles and Moore, 1921: 8 (as Ideobsiidae [sic]); Chamberlin, 1930: 22; Chamberlin, 1931a: 217; Beier, 1932a: 141; Roewer, 1937: 250; Hoff, 1956b: 4; Morikawa, 1960: 119.

Genus Alocobisium Beier

Alocobisium Beier, 1952a: 98-99; Harvey, 1991a: 417.

 

Type species: Alocobisium malaccense Beier, 1952a, by original designation.

Distribution: ASIA: widespread from India to Solomon Islands.

Alocobisium himalaiense Beier

Alocobisium himalaiense Beier, 1976e: 97-98, fig. 3; Harvey, 1991a: 417.

 

Type locality: Phuntsholing, Chhukha, Bhutan.

Distribution: Bhutan.

Alocobisium malaccense Beier

Alocobisium malaccense Beier, 1952a: 99-100, figs 3a-d; Harvey, 1991a: 417.

 

Type locality: Sungei Buloh Leper Settlement, Selangor, Malaysia.

Distribution: Malaysia (mainland).

Alocobisium ocellatum Beier

Alocobisium ocellatum Beier, 1978d: 231-232, fig. 1; Harvey, 1991a: 417; Schawaller, 1994: 742.

 

Type locality: Kaziranga, Assam, India.

Distribution: India; Thailand.

Alocobisium philippinense Beier

Alocobisium philippinense Beier, 1966b: 342; Tenorio and Muchmore, 1982: 383; Harvey, 1991a: 417.

 

Type locality: Tarumpitao Point, Palawan, Philippines.

Distribution: Philippines.

Alocobisium rahmi Beier

Alocobisium rahmi Beier, 1976e: 98-99, fig. 4; Schawaller, 1983a: 108-110, figs 8-11; Schawaller, 1987c: 209; Harvey, 1991a: 418; Schawaller, 1991b: 779, fig. 1.

 

Type locality: Phuntsholing, Chhukha, Bhutan.

Distribution: Bhutan; Nepal.

Alocobisium solomonense Morikawa

Alocobisium solomonense Morikawa, 1963: 3-4, figs 1d-f; Beier, 1964f: 593-594; Beier, 1965g: 762; Beier, 1966d: 138; Beier, 1970g: 317; Muchmore, 1982c: 218; Harvey, 1991a: 418.

 

Type locality: Honiara (as Honiala [sic]), Guadalcanal Island, Guadalcanal, Solomon Islands.

Distribution: Solomon Islands.

Genus Ideobisium Balzan

Ideobisium Balzan, 1892: 539-540; With, 1906: 81; Chamberlin, 1930: 36; Beier, 1932a: 157; Beier, 1976f: 210; Muchmore, 1982c: 194-195; Harvey, 1991a: 419-420.

Ideobisium (Ideobisium) Balzan: Balzan, 1892: 542.

 

Type species: Ideobisium (Ideobisium) crassimanum Balzan, 1892, by subsequent designation of Chamberlin, 1930: 36.

Distribution: Circum-tropical.

Ideobisium antipodum (Simon)

Obisium antipodum Simon, 1880a: 174-175; Rainbow, 1897: 108.

Ideobisium (Ideobisium) antipodanum (Simon): Balzan, 1892: 543 (as Ideobisium (Ideobisium) antipodano [sic]).

Ideobisium (?) antipodum (Simon): Beier, 1932a: 160; Chamberlin, 1934b: 4; Roewer, 1937: 252.

Ideobisium antipodum (Simon): Beier, 1968a: 763-764, fig. 4; Muchmore, 1982c: 206; Harvey, 1991a: 420.

 

Type locality: Nouméa, New Caledonia.

Distribution: New Caledonia; Tuvalu.

Ideobisium balzanii With

Ideobisium balzanii With, 1905: 131-135, plate 10 figs 2a-h; Chamberlin, 1930: 37; Chamberlin, 1931a: fig. 35o; Beier, 1932a: 158 (as Ideobisium balzani); Roewer, 1937: 252 (as Ideobisium balzani); Hoff, 1945e: 1; Muchmore, 1982c: 197-198, fig. 10; Heurtault and Rebière, 1983: 598-600, figs 16-21 (as Ideobisium balzani); Harvey, 1991a: 420; Judson, 1997a: 10.

 

Type locality: St Vincent, St Vincent and the Grenadines.

Distribution: Dominica; Guadeloupe; St Vincent and the Grenadines.

Ideobisium chapmani Muchmore

Ideobisium chapmani Muchmore, 1982c: 198-200, fig. 11; Harvey, 1991a: 420; Muchmore, 1993a: 87.

 

Type locality: Camburales Cave, 10 km E. of Curimagua, Serrania de San Luis, Falcón, Venezuela.

Distribution: Venezuela.

Ideobisium crassimanum Balzan

Ideobisium (Ideobisium) crassimanum Balzan, 1892: 542-543, figs 33, 33a.

Ideobisium crassimanum Balzan: Hansen, 1894: figs 5-7; Chamberlin, 1930: 37; Beier, 1932a: 158, fig. 190; Roewer, 1937: 252; Vachon, 1952a: fig. 16; Beier, 1977c: 100; Beier, 1976d: 46; Muchmore, 1982c: 195-197, figs 2-9; Harvey, 1991a: 420; Muchmore, 1993a: 87.

 

Type locality: Caracas, Distrito Federal, Venezuela.

Distribution: Dominican Republic; Ecuador; Trinidad and Tobago; Venezuela.

Ideobisium ecuadorense Muchmore

Ideobisium ecuadorense Muchmore, 1982c: 201-202, fig. 13; Harvey, 1991a: 420.

 

Type locality: Cueva de los Tayos, Cordillera el Condor, Morona-Santiago, Ecuador.

Distribution: Ecuador.

Ideobisium gracile Balzan

Ideobisium (Ideoroncus) gracilis Balzan, 1892: 540-541, figs 31, 31a.

Ideoroncus gracilis (Balzan): Chamberlin, 1930: 44.

Ideobisium gracile Balzan: Beier, 1932a: 158-159; Roewer, 1937: 252; Harvey, 1991a: 420.

Ideobisium (?) gracile Balzan: Muchmore, 1982c: 206; Muchmore, 1993a: 88.

 

Type locality: San Estéban, Venezuela.

Distribution: Venezuela.

Ideobisium peckorum Muchmore

Ideobisium peckorum Muchmore, 1982c: 200-201, fig. 12; Mahnert, 1985c: 219; Mahnert and Adis, 1986: 214; Harvey, 1991a: 420-421; Muchmore, 1993a: 87; Mahnert and Adis, 2002: 379; Ceballos and Florez, 2007: 50.

 

Type locality: 7 km N. of Leticia, Amazonas, Colombia.

Distribution: Brazil (Amazonas); Colombia.

Ideobisium peregrinum Chamberlin

Ideobisium peregrinum Chamberlin, 1930: 37; Chamberlin, 1931a: figs 11r, 35f-h, 40s, 43j-k, 50c; Beier, 1932a: 159; Chamberlin, 1934b: 4; Roewer, 1937: 252; Beier, 1948c: 537; Beier, 1967b: 290; Beier, 1969e: 414; Wise, 1970: 234; Beier, 1976f: 210; Muchmore, 1982c: 205-206; Harvey, 1991a: 421; Judson, 1997a: 33.

 

Type locality: Days Bay, Wellington, Wellington, North Island, New Zealand.

Distribution: New Zealand.

Ideobisium puertoricense Muchmore

Ideobisium puertoricense Muchmore, 1982c: 202-203, figs 14-15; Harvey, 1991a: 421; Harvey and Volschenk, 2007b: 369, figs 1-4.

 

Type locality: Sierra de Luquillo (as Luquillo Mountains), El Yunque, Fajardo, Puerto Rico.

Distribution: Dominican Republic; Puerto Rico.

Ideobisium puertoricense cavicola Muchmore

Ideobisium puertoricense cavicola Muchmore, 1982c: 203-204 Ideobisium puertoricense cavicolum [sic]); Harvey, 1991a: 421.

 

Type locality: Aguas Buenas Cave, Aguas Buenas, Puerto Rico.

Distribution: Puerto Rico.

Ideobisium puertoricense puertoricense Muchmore

 

Distribution: Dominican Republic; Puerto Rico.

Ideobisium schusteri Mahnert

Ideobisium schusteri Mahnert, 1985c: 220, figs 5-11; Mahnert and Adis, 1986: 214; Harvey, 1991a: 421; Adis and Mahnert, 1993: 435, figs 2-3, 5-6; Mahnert and Adis, 2002: 379; Adis, Bonaldo, Brescovit, Bertani, Cokendolpher, Condé, Kury, Lourenço, Mahnert, Pinto-da-Rocha, Platnick, Reddell, Rheims, Rocha, Rowland, Weygoldt, and Woas, 2002: 5; Aguiar, Gualberto and Franklin, 2006: 795, 796, etc.

 

Type locality: Río Tarumã-Mirím, Manaus, Amazonas, Brazil.

Distribution: Brazil (Amazonas).

Ideobisium trifidum (Stecker)

Obisium trifidum Stecker, 1875b: 523, plate 4 figs 5-8; With, 1906: 77.

Ideobisium (?) trifidum (Stecker): Beier, 1932a: 160; Roewer, 1937: 252.

Genus ? trifidum Stecker: Muchmore, 1982c: 207.

Ideobisium trifidum (Stecker): Harvey, 1991a: 421.

 

Type locality: India.

Distribution: India.

Ideobisium yunquense Muchmore

Ideobisium yunquense Muchmore, 1982c: 204-205, fig. 16; Harvey, 1991a: 421.

 

Type locality: Monte Britton, El Yunque, Rio Grande, Puerto Rico.

Distribution: Puerto Rico.

Genus Ideoblothrus Balzan

Ideobisium (Ideoblothrus) Balzan, 1892: 541.

Pachychitra Chamberlin, 1938b: 111; Hoff, 1945e: 1; Hoff, 1964a: 7-8; Muchmore, 1979a: 195 (synonymised by Muchmore, 1982c: 207).

Ideoblothrus Balzan: Muchmore, 1982c: 207; Harvey, 1991a: 421; Harvey, 1991e: 495; Harvey and Edward, 2007: 449-450.

 

Type species: of Ideobisium (Ideoblothrus): Ideobisium (Ideoblothrus) similis Balzan, 1892, by subsequent designation of Muchmore, 1982c: 207.

of Pachychitra: Pachychitra maya Chamberlin, 1938b, by original designation.

Distribution: Circum-tropical.

Ideoblothrus amazonicus (Mahnert)

Ideobisium amazonicum Mahnert, 1979d: 743-744, figs 48-52.

Ideoblothrus amazonicus (Mahnert): Muchmore, 1982c: 215; Mahnert and Adis, 1986: 214; Harvey, 1991a: 421; Harvey, 1992c: figs 113-116; Mahnert and Adis, 2002: 379.

 

Type locality: Rio Demini (as Demeni), Amazonas, Brazil.

Distribution: Brazil (Amazonas).

Ideoblothrus baloghi (Mahnert)

Ideobisium baloghi Mahnert, 1978f: 90-92, figs 48-50.

Ideoblothrus baloghi (Mahnert): Muchmore, 1982c: 216; Harvey, 1991a: 422.

 

Type locality: Bangu forest, Meya, Kindamba, Pool, Congo.

Distribution: Congo.

Ideoblothrus bipectinatus (Daday)

Ideobisium bipectinatum Daday, 1897: 478-479, figs 7, 14, 15; With, 1906: 87-88; Ellingsen, 1910a: 395; Beier, 1932a: 160, fig. 192; Chamberlin, 1934b: 4; Roewer, 1937: 252; Morikawa, 1963: 4, figs 2a-c; Beier, 1965g: 761-762; Beier, 1967c: 321; Beier, 1982: 43; Hammen, 1983b: 16, 17, 19, 29.

Ideoblothrus bipectinatus (Daday): Muchmore, 1982c: 216; Harvey, 1991a: 422.

 

Type locality: Madang (as Friedrich-Wilhelmshafen), Madang, Papua New Guinea.

Distribution: Indonesia (West Papua); Papua New Guinea.

Ideoblothrus brasiliensis (Mahnert)

Ideobisium brasiliense Mahnert, 1979d: 747-750, figs 59-64; Adis, Junk and Penny, 1987: 488.

Ideoblothrus brasiliensis (Mahnert): Muchmore, 1982c: 215; Mahnert, 1985b: 76; Mahnert and Adis, 1986: 214; Mahnert, Adis and Bührnheim, 1987: fig. 7b; Harvey, 1991a: 422; Mahnert and Adis, 2002: 379, fig. 7b.

 

Type locality: Belém do Pará, Pará, Brazil.

Distribution: Brazil (Amazonas, Pará).

Ideoblothrus caecus (Mahnert)

Ideobisium caecum Mahnert, 1979d: 745-747, figs 53-58.

Ideoblothrus caecus (Mahnert): Muchmore, 1982c: 215; Mahnert and Adis, 1986: 214; Mahnert, Adis and Bührnheim, 1987: fig. 9; Harvey, 1991a: 422; Mahnert and Adis, 2002: 379, fig. 9.

 

Type locality: Santarém, Pará, Brazil.

Distribution: Brazil (Amazonas, Pará).

Ideoblothrus carinatus (Hoff)

Pachychitra carinata Hoff, 1964a: 13-14, figs 5-6.

Ideoblothrus carinatus (Hoff): Muchmore, 1982c: 215; Harvey, 1991a: 422.

 

Type locality: 5 miles S. of Hardwar Gap, Parish of St Andrew, Jamaica.

Distribution: Jamaica.

Ideoblothrus ceylonicus (Beier)

Ideobisium ceylonicum Beier, 1973d: 42-43, fig. 4.

Ideoblothrus ceylonicus (Beier): Harvey, 1991a: 422.

 

Type locality: Andapolakanda, 3 miles NE. of Melsiripura, North-West Province, Sri Lanka.

Distribution: Sri Lanka.

Ideoblothrus colombiae Muchmore

Ideoblothrus colombiae Muchmore, 1982c: 211-212, figs 21-22; Harvey, 1991a: 422; Muchmore, 1993a: 88; Ceballos and Florez, 2007: 50.

 

Type locality: between San Pedro and San Javier, Sierra Nevada de Santa Marta, Magdalena, Colombia.

Distribution: Colombia.

Ideoblothrus costaricensis (Beier)

Ideobisium costaricense Beier, 1931c: 302-303, fig. 2; Beier, 1932a: 159, figs 189, 191; Roewer, 1937: 252; Beier, 1977c: 100.

Ideobisium simile Balzan: Beier, 1974c: 101 (in part).

Ideoblothrus costaricensis (Beier): Muchmore, 1982c: 213; Harvey, 1991a: 422-423.

 

Type locality: Tuis, Cartago, Costa Rica.

Distribution: Costa Rica; Ecuador.

Ideoblothrus curazavius (Wagenaar-Hummelinck)

Pachychitra curazavia Wagenaar-Hummelinck, 1948: 63-71, figs 22a-e, 23a-g, 24a-f, 25a-f, 26h.

Ideoblothrus curazavius (Wagenaar-Hummelinck): Muchmore, 1982c: 214; Harvey, 1991a: 423; Muchmore, 1993a: 88.

 

Type locality: Seroe Christoffel, Curaçao, Netherlands Antilles.

Distribution: Netherlands Antilles (Curaçao).

Ideoblothrus descartes Harvey and Edward

Ideoblothrus descartes Harvey and Edward, 2007: 465-468, figs 36-41.

 

Type locality: Descartes Island, Western Australia, Australia.

Distribution: Australia (Western Australia).

Ideoblothrus fenestratus (Beier)

Ideobisium fenestratum Beier, 1955m: 3-4, figs 3-5; Weidner, 1959: 115.

Ideoblothrus fenestratus (Beier): Muchmore, 1982c: 214; Harvey, 1991a: 423; Mahnert and Adis, 2002: 379.

 

Type locality: Sivia, Ayacucho, Peru.

Distribution: Peru.

Ideoblothrus floridensis (Muchmore)

Pachychitra floridensis Muchmore, 1979a: 195-197, figs 1-6.

Ideoblothrus floridensis (Muchmore): Muchmore, 1982c: 215; Muchmore, 1990a: 513, fig. 18.39; Harvey, 1991a: 423.

 

Type locality: Watson Hammock, Big Pine Key, Monroe County, Florida, U.S.A.

Distribution: U.S.A. (Florida).

Ideoblothrus godfreyi (Ellingsen)

Ideobisium (Ideoblothrus) godfreyi Ellingsen, 1912b: 117-118.

Gymnobisium godfreyi (Ellingsen): Beier, 1932a: 162; Roewer, 1937: 253.

Ideobisium godfreyi (Ellingsen): Beier, 1947b: 290.

Ideoblothrus godfreyi (Ellingsen): Muchmore, 1982c: 215; Harvey, 1991a: 423.

 

Type locality: Frankfort Hill, near King William's Town, Eastern Cape, South Africa.

Distribution: South Africa.

Ideoblothrus grandis (Muchmore)

Pachychitra grandis Muchmore, 1972g: 266, figs 6-7; Muchmore, 1977: 70; Reddell, 1981: 122-123, fig. 20; Ceballos, 2004: 428.

Ideoblothrus grandis (Muchmore): Muchmore, 1982c: 214; Harvey, 1991a: 423; Muchmore, 1991b: 156; Villegas-Guzman, 2006: 133.

 

Type locality: Cueva del Tío Ticho, 1 mile S. of Comitan, Chiapas, Mexico.

Distribution: Mexico.

Ideoblothrus holmi (Beier)

Ideobisium holmi Beier, 1955c: 534, fig. 5; Beier, 1959d: 23; Beier, 1972a: 7.

Ideoblothrus holmi (Beier): Muchmore, 1982c: 216; Harvey, 1991a: 423.

 

Type locality: Bundibugyo, Ruwenzori, Uganda.

Distribution: Democratic Republic of Congo (Zaire); Uganda.

Ideoblothrus insularum (Hoff)

Pachychitra insularum Hoff, 1945e: 1-4, figs 1-5; Wagenaar-Hummelinck, 1948: 73-75, figs 29a-l; Hoff, 1964a: 8-9.

Ideoblothrus insularum (Hoff): Muchmore, 1982c: 214; Harvey, 1991a: 423.

 

Type locality: Desecheo Island, Puerto Rico.

Distribution: Jamaica; Puerto Rico.

Ideoblothrus kochalkai Muchmore

Ideoblothrus kochalkai Muchmore, 1982c: 210, figs 18-20; Harvey, 1991a: 424; Muchmore, 1993a: 88.

 

Type locality: Casa Antonio, Cuchilla Cebolleta, Sierra Nevada de Santa Marta, Magdalena, Colombia.

Distribution: Colombia.

Ideoblothrus leleupi (Beier)

Ideobisium leleupi Beier, 1959d: 23-25, fig. 10.

Ideoblothrus leleupi (Beier): Muchmore, 1982c: 216; Harvey, 1991a: 424; Beron, 2002: 35.

 

Type locality: Lubero, Haut-Zaire, Democratic Republic of Congo (Zaire).

Distribution: Democratic Republic of Congo (Zaire).

Ideoblothrus lepesmei (Vachon)

Ideobisium lepesmei Vachon, 1941e: 32; Vachon, 1952a: 24-27, figs 10-11, 13-14.

Ideoblothrus lepesmei (Vachon): Muchmore, 1982c: 216; Harvey, 1991a: 424.

 

Type locality: Sassandra, Côte d’Ivoire.

Distribution: Côte d’Ivoire; Guinea.

Ideoblothrus levipalpus Mahnert

Ideoblothrus levipalpus Mahnert, 1985c: 221, figs 12-13; Mahnert and Adis, 1986: 214; Harvey, 1991a: 424; Mahnert and Adis, 2002: 379.

 

Type locality: S. of Rio Negro, Amazonas, Brazil.

Distribution: Brazil (Amazonas).

Ideoblothrus maya (Chamberlin)

Pachychitra maya Chamberlin, 1938b: 111-113, figs 1a-f; Roewer, 1940: 345, fig. 261; Wagenaar-Hummelinck, 1948: 71-73, figs 26a-g, 27a-c, 28a-f; Nicholas, 1962: 181; Reddell, 1971a: 27; Muchmore, 1977: 70.

Ideoblothrus maya (Chamberlin): Muchmore, 1982c: 213; Harvey, 1991a: 424; Muchmore, 1991b: 156; Ceballos, 2004: 428.

 

Type locality: cave near Oxkutzcab, Yucatán, Mexico.

Distribution: Mexico.

Ideoblothrus mexicanus (Muchmore)

Pachychitra mexicana Muchmore, 1972g: 262-264, figs 1-3.

Ideoblothrus mexicanus (Muchmore): Muchmore, 1982c: 214; Harvey, 1991a: 424; Ceballos, 2004: 428.

 

Type locality: Rancho del Cielo, 6 miles NW. of Gomez Farais, Tamaulipas, Mexico.

Distribution: Mexico.

Ideoblothrus muchmorei Heurtault

Ideoblothrus muchmorei Heurtault, 1983: 24-26, figs 53-58; Harvey, 1991a: 424.

 

Type locality: Bandama, Côte d’Ivoire.

Distribution: Côte d’Ivoire.

Ideoblothrus milikapiti Harvey and Edward

Ideoblothrus milikapiti Harvey and Edward, 2007: 468-470, figs 42-45.

 

Type locality: Milikapiti, Northern Territory, Australia.

Distribution: Australia (Northern Territory).

Ideoblothrus nesotymbus Harvey and Edward

Ideoblothrus nesotymbus Harvey and Edward, 2007: 457-462, figs 4, 22-28.

 

Type locality: 5.8 km NW. of Chevron Texaco Camp, borehole S2, Barrow Island, Western Australia, Australia.

Distribution: Australia (Western Australia).

Ideoblothrus occidentalis (Beier)

Ideobisium occidentale Beier, 1959d: 25-26, fig. 11.

Ideoblothrus occidentalis (Beier): Muchmore, 1982c: 216; Harvey, 1991a: 424.

 

Type locality: Inkisi-Kisantu (as d'Inkisi, Kisantu Mission), Zaire Central, Democratic Republic of Congo (Zaire).

Distribution: Democratic Republic of Congo (Zaire).

Ideoblothrus palauensis (Beier)

Ideobisium palauense Beier, 1957d: 13-14, fig. 4a.

Ideoblothrus palauensis (Beier): Muchmore, 1982c: 217; Harvey, 1991a: 424-425.

 

Type locality: East Ngatpang, Babelthuap, Palau.

Distribution: Palau.

Ideoblothrus papillon Harvey

Ideoblothrus papillon Harvey, 1991e: 498-500, figs 34-41; Harvey, Gray, Hunt and Lee, 1993: 131, 134; Harvey and Edward, 2007: 450.

 

Type locality: Papillon Cave, C-15, Cape Range peninsula, Western Australia, Australia.

Distribution: Australia (Western Australia).

Ideoblothrus paraensis Mahnert

Ideoblothrus paraensis Mahnert, 1985b: 76-78, figs 1-6; Mahnert and Adis, 1986: 214; Harvey, 1991a: 425; Mahnert and Adis, 2002: 379.

 

Type locality: Paricatuba, Benevides, Pará, Brazil.

Distribution: Brazil (Pará).

Ideoblothrus pisolitus Harvey and Edward

Ideoblothrus pisolitus Harvey and Edward, 2007: 452-455, figs 3, 8-21.

 

Type locality: Mesa B, 38.1 km W. of Pannawonica, Western Australia, Australia.

Distribution: Australia (Western Australia).

Ideoblothrus pugil (Beier)

Ideobisium pugil Beier, 1964f: 593, fig. 1; Beier, 1965g: 762; Beier, 1966d: 137; Beier, 1970g: 316.

Ideoblothrus pugil (Beier): Harvey, 1991a: 425; Judson, 1997a: 35-36.

 

Type locality: Mt Austen, Guadalcanal Island, Guadalcanal, Solomon Islands.

Distribution: Solomon Islands.

Ideoblothrus pugil pugil (Beier)

Ideobisium pugil pugil Beier: Petersen, 1968: 119.

Ideoblothrus pugil pugil (Beier): Muchmore, 1982c: 217.

 

Distribution: Solomon Islands.

Ideoblothrus pugil robustus (Beier)

Ideobisium pugil robustum Beier, 1966d: 137, fig. 3.

Ideoblothrus pugil robustus (Beier): Muchmore, 1982c: 217; Harvey, 1991a: 425; Judson, 1997a: 37.

 

Type locality: Nila, Shortland Islands, Western, Solomon Islands.

Distribution: Solomon Islands.

Ideoblothrus pygmaeus (Hoff)

Pachychitra pygmaea Hoff, 1964a: 9-11, figs 1-2.

Ideoblothrus pygmaeus (Hoff): Muchmore, 1982c: 214; Heurtault and Rebière, 1983: 596-598, figs 11-15; Harvey, 1991a: 425.

 

Type locality: Dolphin Head, 5 miles S. of Lucea, Hanover Parish, Jamaica.

Distribution: Jamaica; Martinique.

Ideoblothrus seychellesensis (Chamberlin)

Ideobisium seychellesensis Chamberlin, 1930: 38, figs 1x, 1dd, 2cc; Chamberlin, 1931a: figs 13i, 16l, 17g; Beier, 1932a: 160; Roewer, 1937: 252.

Ideoblothrus seychellesensis (Chamberlin): Muchmore, 1982c: 212-213, figs 23-24; Harvey, 1991a: 425; Judson, 1997a: 39-40; Harvey, 2001b: 691.

 

Type locality: Félicité Island, Seychelles.

Distribution: Seychelles.

Ideoblothrus similis (Balzan)

Ideobisium (Ideoblothrus) simile Balzan, 1892: 541-542, figs 32, 32a (as Ideobisium (Ideoblothrus) similis).

Ideobisium simile Balzan: Beier, 1932a: 159; Ellingsen, 1909b: 219; Roewer, 1937: 252; Vachon, 1952a: figs 12, 15; Beier, 1974c: 101 (in part; see Ideoblothrus costaricensis (Beier)); Beier, 1977c: 100-101; Hounsome, 1980: 85; Baert, Maelfait and Desender, 1995: 18.

Ideoblothrus similis (Balzan): Muchmore, 1982c: 208-210, fig. 17; Harvey, 1991a: 425; Muchmore, 1993a: 87.

 

Type locality: Petare, Miranda, Venezuela.

Distribution: Cayman Islands; Ecuador (Galapagos Islands); Mexico; Venezuela.

Ideoblothrus tenuis Mahnert

Ideoblothrus tenuis Mahnert, 1985c: 221-222, figs 14-18; Mahnert and Adis, 1986: 214; Adis and Mahnert, 1990b: 13, figs 2-3; Harvey, 1991a: 425; Adis and Mahnert, 1993: 435, fig. 2; Mahnert and Adis, 2002: 379; Adis, Bonaldo, Brescovit, Bertani, Cokendolpher, Condé, Kury, Lourenço, Mahnert, Pinto-da-Rocha, Platnick, Reddell, Rheims, Rocha, Rowland, Weygoldt, and Woas, 2002: 5; Aguiar, Gualberto and Franklin, 2006: 795, 796, etc.

 

Type locality: Reserva Florestal Adolpho Ducke, 26 km N. of Manaus, on Manaus-Itacoatiara Highway, Amazonas, Brazil.

Distribution: Brazil (Amazonas).

Ideoblothrus truncatus (Hoff)

Pachychitra truncata Hoff, 1964a: 11-13, figs 3-4.

Ideoblothrus truncatus (Hoff): Muchmore, 1982c: 215; Harvey, 1991a: 426.

 

Type locality: Maggotty Falls, S. of Maggotty, Parish of St Elizabeth, Jamaica.

Distribution: Jamaica.

Ideoblothrus vampirorum Muchmore

Pachychitra similis Muchmore, 1972g: 264-265, figs 4-5; Reddell and Elliott, 1973b: 183 (junior secondary homonym of Ideoblothrus similis (Balzan)).

Ideoblothrus vampirorum Muchmore, 1982c: 214; Harvey, 1991a: 426; Ceballos, 2004: 428 (replacement name for Pachychitra similis Muchmore).

 

Type locality: Cueva de los Vampiros, 6 miles NNE. of Chamal, Tamaulipas, Mexico.

Distribution: Mexico.

Ideoblothrus westi Harvey and Edward

Ideoblothrus westi Harvey and Edward, 2007: 462-465, figs 29-35.

 

Type locality: Fortesque River, borehole B7, Western Australia, Australia.

Distribution: Australia (Western Australia).

Ideoblothrus woodi Harvey

Ideoblothrus woodi Harvey, 1991e: 497-498, figs 27-33, 40-41; Harvey, Gray, Hunt and Lee, 1993: 131, 134; Harvey and Edward, 2007: 450-452, figs 5, 6.

 

Type locality: Cave C-167, Cape Range peninsula, Western Australia, Australia.

Distribution: Australia (Western Australia).

Ideoblothrus zicsii (Mahnert)

Ideobisium zicsii Mahnert, 1978f: 92-93, figs 51-55.

Ideoblothrus zicsii (Mahnert): Muchmore, 1982c: 216; Harvey, 1991a: 426.

 

Type locality: Sibiti, Lékoumou, Congo.

Distribution: Congo.

Genus Microblothrus Mahnert

Microblothrus Mahnert, 1985c: 222; Harvey, 1991a: 426.

 

Type species: Microblothrus tridens Mahnert, 1985c, by original designation.

Distribution: SOUTH AMERICA: Brazil.

Microblothrus tridens Mahnert

Microblothrus tridens Mahnert, 1985c: 222-223, figs 19-26; Mahnert and Adis, 1986: 214; Adis and Mahnert, 1990b: 13, figs 2-3; Harvey, 1991a: 426; Adis and Mahnert, 1993: 435, figs 2-3, 5-6; Höfer and Beck, 1995: 398; Mahnert and Adis, 2002: 380; Adis, Bonaldo, Brescovit, Bertani, Cokendolpher, Condé, Kury, Lourenço, Mahnert, Pinto-da-Rocha, Platnick, Reddell, Rheims, Rocha, Rowland, Weygoldt, and Woas, 2002: 5; Aguiar, Gualberto and Franklin, 2006: 795, 796, etc.

 

Type locality: Reserva Florestal Adolpho Ducke, 26 km N. of Manaus, on Manaus-Itacoatiara Highway, Amazonas, Brazil.

Distribution: Brazil (Amazonas).

Genus Nannobisium Beier

Nannobisium Beier, 1931c: 303; Beier, 1932a: 161; Harvey, 1991a: 427.

Vescichitra Hoff, 1964a: 15 (synonymised by Mahnert, 1979d: 752).

 

Type species: of Nannobisium: Nannobisium liberiense Beier, 1931c, by original designation.

of Vescichitra: Vescichitra mollis Hoff, 1964a, by original designation.

Distribution: AFRICA: west Africa. SOUTH AMERICA: Brazil; Jamaica.

Nannobisium beieri Mahnert

Nannobisium beieri Mahnert, 1979d: 751-752, figs 65-69; Mahnert, 1985b: 76; Mahnert and Adis, 1986: 214; Mahnert, Adis and Bührnheim, 1987: fig. 8; Harvey, 1991a: 427; Mahnert, 2001: 103; Mahnert and Adis, 2002: 380, fig. 8.

 

Type locality: Santarém, Pará, Brazil.

Distribution: Brazil (Pará).

Nannobisium liberiense Beier

Nannobisium liberiense Beier, 1931c: 303-304, fig. 3; Beier, 1932a: 161, figs 193-194; Roewer, 1936: fig. 75; Roewer, 1937: 252; Vachon, 1941e: 31; Mahnert, 1974d: 851; Muchmore, 1982c: 218; Harvey, 1991a: 427.

 

Type locality: Bolahun, Lofa, Liberia.

Distribution: Côte d’Ivoire; Liberia; Togo.

Nannobisium mollis (Hoff)

Vescichitra mollis Hoff, 1964a: 15-18, figs 7-8.

Nannobisium mollis (Hoff): Mahnert, 1979d: 752; Harvey, 1991a: 427.

 

Type locality: 1 miles NW. of Ferry, Parish of St Andrew, Jamaica.

Distribution: Jamaica.

Subfamily SYARININAE Chamberlin

Syarininae Chamberlin, 1930: 39; Chamberlin, 1931a: 219; Beier, 1932a: 163; Chamberlin, 1938b: 109-110; Hoff, 1956b: 10.

Genus Anysrius Harvey

Anysrius Harvey, 1998: 430-431.

 

Type species: Anysrius chamberlini Harvey, 1998, by original designation.

Distribution: AUSTRALASIA: Tasmania.

Anysrius brochus Harvey

Anysrius brochus Harvey, 1998: 434-437, figs 19-31.

 

Type locality: 'Chatlee Road' site, Salmon River Forestry area, Tasmania, Australia.

Distribution: Australia (Tasmania).

Anysrius chamberlini Harvey

Undescribed genus and species: Harvey, 1990d: 158-159, fig. 4; Harvey 1996a: 258.

Anysrius chamberlini Harvey, 1998: 431-434, figs 1-18; Harvey and Volschenk, 2007b: 369, figs 1-4.

 

Type locality: Frodshams Pass, Tasmania, Australia.

Distribution: Australia (Tasmania).

Genus Syarinus Chamberlin

Syarinus Chamberlin, 1925a: 329; Chamberlin, 1930: 39; Beier, 1932a: 163; Hoff, 1956b: 10; Harvey, 1991a: 429; Harvey, 1998: 437-439.

 

Type species: Ideoroncus obscurus Banks, 1893, by original designation.

Distribution: NORTH AMERICA: widespread in U.S.A. and Canada. EUROPE: northern and central Europe.

Syarinus enhuycki Muchmore

Syarinus enhuycki Muchmore, 1968e: 112-115, figs 1-3; Nelson, 1975: 281-282, figs 25-28; Harvey, 1990d: 158; Harvey, 1991a: 429; Snider and Nelson, 1991: 260; Harvey, 1998: 438; Buddle, 2005: 15; Watermolen, 2007: 3.

 

Type locality: E.N. Huyck Preserve, Rensselaerville, Albany County, New York, U.S.A.

Distribution: Canada (Ontario); U.S.A. (Michigan, New Hampshire, New York, Pennsylvania, Wisconsin).

Syarinus granulatus Chamberlin

Syarinus granulatus Chamberlin, 1930: 39-40, fig. 2h; Chamberlin, 1931a: figs 9e, 11v, 13m, 15g, 16j, 17n, 28n, 36a; Beier, 1932a: 164; Roewer, 1937: 254; Hoff and Bolsterli, 1956: 163; Hoff, 1956b: 16-20, fig. 10; Hoff, 1958: 13; Hoff, 1959b: 4, etc.; Lindroth and Ball, 1969: ? (not seen); Harvey, 1991a: 429; Harvey, 1998: 438.

 

Type locality: Englemann Canyon, Manitou, El Paso County, Colorado, U.S.A.

Distribution: U.S.A. (Alaska, Colorado, New Mexico).

Syarinus honestus Hoff

Syarinus honestus Hoff, 1956b: 14-16, figs 8-9; Hoff, 1958: 14; Hoff, 1959b: 4, etc.; Harvey, 1991a: 429.

 

Type locality: Santa Fe ski area, NE. of Santa Fe, Santa Fe County, New Mexico, U.S.A.

Distribution: U.S.A. (New Mexico).

Syarinus obscurus (Banks)

Ideoroncus obscurus Banks, 1893: 66-67; Banks, 1895a: 11; Banks, 1904b: 364; Coolidge, 1908: 113; Banks, 1911: 639; Moles, 1914c: 196-197; Banks, 1916: 72; Moles and Moore, 1921: 8.

Ideobisium obscurum (Banks): Ellingsen, 1908e: 163.

Syarinus obscurus (Banks): Chamberlin, 1925a: 330; Chamberlin, 1930: 39; Chamberlin, 1931a: figs 9d, 15f, 17l-m, 19e, 25k, 43s-t, 47d; Beier, 1932a: 164; Roewer, 1937: 253; Hoff, 1956b: 10-14, figs 6-7; Hoff, 1958: 13; Hoff, 1959b: 4, etc.; Muchmore, 1971a: 82; Knowlton, 1972: 2; Muchmore, 1973g: 122; Knowlton, 1974: 3; Harvey, 1991a: 429; Scudder, 1994: 18; Harvey, 1998: 438, figs 32-33; Poinar, Ćurčić and Cokendolpher, 1998: 89; Buddle, 2005: 14, 15.

 

Type locality: Olympia, Thurston County, Washington, U.S.A.

Distribution: Canada (British Columbia, Saskatchewan, Yukon Territory); U.S.A. (California, Montana, New Mexico, Utah, Washington, Wyoming).

Syarinus palmeni Kaisila

Syarinus palmeni Kaisila, 1964: 52-54, figs 1a-e (as Syarinus palméni); Harvey, 1991a: 430; Buddle, 2005: 15.

 

Type locality: Holyrood, Newfoundland, Canada.

Distribution: Canada (Newfoundland).

Syarinus strandi (Ellingsen)

Ideobisium (Ideoblothrus) strandi Ellingsen, 1901a: 88-89; Ellingsen, 1903: 12-13.

Ideobisium strandi Ellingsen: Ellingsen, 1910c: 348.

Microcreagris strandi (Ellingsen): Beier, 1932a: 155; Roewer, 1937: 251; Kaisila, 1949b: 79-80, fig. 2, map 4; Vachon, 1954d: 591; Meinertz, 1962: 56, fig. 33b; Beier, 1963b: 211; Beier, 1970f: 321; Klausen, 1975: 64.

Syarinus strandi (Ellingsen): Mahnert, 1976c: 206-209, figs 20-25; Schawaller, 1987b: 289-292, figs 1-10; Harvey, 1991a: 430; Gärdenfors and Wilander, 1992: 29; Platen, Blick, Bliss, Drogla, Malten, Martens, Sacher and Wunderlich, 1995: 54; Drogla and Blick, 1996: 37; Schmarda, 1997: 105-107, figs 1-9; Ducháč, 1998b: 506-508, fig. 1; Ducháč, 1999a: 14; Šťáhlavský and Ducháč, 2001: 48; Drogla and Lippold, 2004: 25, fig. 2.4; Mahnert, 2004: 464; Šťáhlavský and Ducháč, 2005: 83.

 

Type locality: Hallingdal, Buskerud, Norway.

Distribution: Austria; Czech Republic; Germany; Norway; Russia (Leningradskaya Oblast').

 

UNPLACED

Genus Hyarinus Chamberlin

Hyarinus Chamberlin, 1925a: 327-328; Beier, 1932a: 164; Harvey, 1991a: 419.

 

Type species: Hyarinus hesperus Chamberlin, 1925a, by original designation.

Distribution: NORTH AMERICA: U.S.A.

Hyarinus hesperus Chamberlin

Hyarinus hesperus Chamberlin, 1925a: 329, figs a-d; Chamberlin, 1931a: fig. 36b; Beier, 1932a: 165, fig. 196; Vachon, 1934i: 413; Roewer, 1937: 254; Hoff, 1958: 14; Muchmore, 1990a: 513; Harvey, 1991a: 419.

 

Type locality: Santa Barbara, Santa Barbara County, California, U.S.A.

Distribution: U.S.A. (California).

CATALOGUE

 

Family Syarinidae Chamberlin 1930