Pseudoscorpions of the World

Last updated: 19 January 2009

Version 1.2, 2009

CATALOGUE

 

Family Geogarypidae Chamberlin 1930

Family GEOGARYPIDAE Chamberlin

Geogarypinae Chamberlin, 1930: 609; Chamberlin, 1931a: 228; Beier, 1932a: 227; Roewer, 1937: 268-269; Morikawa, 1960: 132; Murthy and Ananthakrishnan, 1977: 104.

Geogarypidae Chamberlin: Harvey, 1986: 754; Harvey, 1991a: 249; Harvey, 1992c: 1419-1420.

Genus Afrogarypus Beier

Afrogarypus Beier, 1931c: 317; Harvey, 1986: 758; Harvey, 1991a: 249.

Geogarypus (Afrogarypus) Beier: Beier, 1932a: 236; Heurtault, 1970b: 1365.

 

Type species: Garypus senegalensis Balzan, 1892, by original designation.

Distribution: AFRICA: widespread.

Afrogarypus basilewskyi (Beier)

Geogarypus (Afrogarypus) basilewskyi Beier, 1962f: 16-18, figs 5-6; Mahnert, 1982d: 318-319, figs 27-28, 54, 57.

Afrogarypus basilewskyi (Beier): Harvey, 1986: 758; Harvey, 1991a: 249; Beron, 2002: 33.

 

Type locality: Ulugu-Ndogo Valley, Uluguru Mountains, Morogoro, Tanzania.

Distribution: Kenya; Tanzania.

Afrogarypus curtus (Beier)

Geogarypus (Afrogarypus) curtus Beier, 1955c: 544, fig. 11; Mahnert, 1982d: 325-326, figs 43, 52, 56.

Afrogarypus curtus (Beier): Harvey, 1986: 758; Harvey, 1991a: 249-250.

 

Type locality: 7 miles N. of Mombasa, Coast, Kenya.

Distribution: Kenya.

Afrogarypus excelsus (Beier)

Geogarypus (Afrogarypus) excelsus Beier, 1964k: 61-62, fig. 22.

Afrogarypus excelsus (Beier): Harvey, 1986: 758; Harvey, 1991a: 250.

 

Type locality: Paradise Kloof, Grahamstown, Eastern Cape, South Africa.

Distribution: South Africa.

Afrogarypus excelsus excellens (Beier)

Geogarypus (Afrogarypus) excelsus excellens Beier, 1964k: 62-63, fig. 23; Beier, 1966k: 461.

Afrogarypus excelsus excellens (Beier): Harvey, 1986: 758; Harvey, 1991a: 250.

 

Type locality: unknown.

Distribution: South Africa.

Afrogarypus excelsus excelsus (Beier)

 

Distribution: South Africa.

Afrogarypus impressus (Tullgren)

Garypus impressus Tullgren, 1907b: 228-229, fig. 7; Ellingsen, 1912b: 86, 107-108.

Geogarypus impressus (Tullgren): Chamberlin, 1930: 609; Roewer, 1936: fig. 77; Vachon, 1949: fig. 204b; Beier, 1964k: 60; Lawrence, 1967: 89; Spaull, 1979: 117.

Geogarypus (Afrogarypus) impressus (Tullgren): Beier, 1932a: 236-237, fig. 265; Roewer, 1937: 270, fig. 222b; Beier, 1947b: 320; Beier, 1955l: 301, fig. 21a; Beier, 1958a: 171; Beier, 1966k: 461.

Afrogarypus impressus (Tullgren): Harvey, 1986: 758; Harvey, 1991a: 250.

 

Type locality: Van Reenen, Free State, South Africa; Amanzimtoti, KwaZulu-Natal, South Africa; and junction of Black and White Umfolozi, KwaZulu-Natal, South Africa.

Distribution: Mozambique; Seychelles (Groupe d'Aldabra); South Africa.

Afrogarypus intermedius (Beier)

Geogarypus (Afrogarypus) intermedius Beier, 1955c: 540-541, fig. 9; Beier, 1955h: 9; Beier, 1959d: 31.

Afrogarypus intermedius (Beier): Harvey, 1986: 758; Harvey, 1991a: 250; Beron, 2002: 33.

 

Type locality: Bundibugyo, Ruwenzori, Uganda.

Distribution: Democratic Republic of Congo (Zaire); Kenya; Rwanda; Uganda.

Afrogarypus intermedius intermedius (Beier)

Geogarypus (Afrogarypus) intermedius intermedius Beier: Mahnert, 1982d: 318.

 

Distribution: Democratic Republic of Congo (Zaire); Kenya; Rwanda; Uganda.

Afrogarypus intermedius nanus (Beier)

Geogarypus (Afrogarypus) intermedius nanus Beier, 1959d: 32-33, fig. 14.

Afrogarypus intermedius nanus (Beier): Harvey, 1986: 758; Harvey, 1991a: 250.

 

Type locality: Kundelungu, Shaba, Democratic Republic of Congo (Zaire).

Distribution: Democratic Republic of Congo (Zaire).

Afrogarypus monticola (Beier)

Geogarypus (Afrogarypus) monticola Beier, 1955c: 541-544, fig. 10; Beier, 1962f: 15-16, figs 3-4; Mahnert, 1982d: 314-318, figs 24-26, 53, 58.

Afrogarypus monticola (Beier): Harvey, 1986: 758; Harvey, 1991a: 250; Beron, 2002: 33.

 

Type locality: Mt Elgon, Western, Kenya.

Distribution: Kenya; Uganda; Tanzania.

Afrogarypus plumatus (Mahnert)

Geogarypus (Afrogarypus) plumatus Mahnert, 1982d: 320-322, figs 29-35, 59.

Afrogarypus plumatus (Mahnert): Harvey, 1986: 758; Harvey, 1991a: 250-251.

 

Type locality: Makadara Forest, Shimba Hills, Coast, Kenya.

Distribution: Kenya.

Afrogarypus pseudocurtus (Mahnert)

Geogarypus (Afrogarypus) pseudocurtus Mahnert, 1982d: 322-325, figs 44-48, 51.

Afrogarypus pseudocurtus (Mahnert): Harvey, 1986: 758; Harvey, 1991a: 251.

 

Type locality: Ngao, Lake Shakababo, Tana River, Coast, Kenya.

Distribution: Kenya.

Afrogarypus senegalensis (Balzan)

Garypus senegalensis Balzan, 1892: 535-536, figs 27, 27a-b; Ellingsen, 1913a: 454.

Geogarypus senegalensis (Balzan): Chamberlin, 1930: 609; Majer, 1974: 90.

Afrogarypus senegalensis (Balzan): Beier, 1931c: 317; Harvey, 1986: 758, fig. 2; Harvey, 1991a: 251.

Geogarypus (Afrogarypus) senegalensis (Balzan): Beier, 1932a: 236, fig. 264; Roewer, 1937: 270; Vachon, 1952a: 27; Beier, 1965b: 371.

Not Garypus senegalensis Balzan: Ellingsen, 1912b: 87, 112-114 (misidentification; see Geogarypus olivaceus (Tullgren)).

 

Type locality: Senegal.

Distribution: Cameroon; Ghana; Guinea; Liberia; Senegal.

Afrogarypus seychellesensis (Beier)

Geogarypus seychellesensis Beier, 1940a: 173-174, fig. 173.

Geogarypus (Afrogarypus) seychellesensis (Beier): Mahnert, 1978d: 878.

Afrogarypus seychellesensis (Beier): Harvey, 1986: 758; Harvey, 1991a: 251; Judson, 1997a: 39-40; Harvey, 2001b: 691.

 

Type locality: Praslin Island, Seychelles.

Distribution: Seychelles.

Afrogarypus stellifer (Mahnert)

Geogarypus (Afrogarypus) stellifer Mahnert, 1982d: 322, figs 36-42, 49, 55.

Afrogarypus stellifer (Beier): Harvey, 1986: 758; Harvey, 1991a: 251.

 

Type locality: Mombasa, Coast, Kenya.

Distribution: Kenya.

Afrogarypus subimpressus (Beier)

Geogarypus (Afrogarypus) subimpressus Beier, 1955l: 302-303, figs 21b, 22.

Geogarypus subimpressus Beier: Beier, 1964k: 60.

Afrogarypus subimpressus (Beier): Harvey, 1986: 758; Harvey, 1991a: 251.

 

Type locality: Cape Point, Cape Peninsula, Western Cape, South Africa.

Distribution: South Africa.

Afrogarypus sulcatus (Beier)

Geogarypus (Afrogarypus) sulcatus Beier, 1955a: 10-12, fig. 5.

Afrogarypus sulcatus (Beier): Harvey, 1986: 758; Harvey, 1991a: 251.

 

Type locality: Mabwe, Shaba, Democratic Republic of Congo (Zaire); Zimbabwe.

Distribution: Democratic Republic of Congo (Zaire).

Afrogarypus sulcatus rhodesiacus (Beier)

Geogarypus (Afrogarypus) sulcatus rhodesiacus Beier, 1964k: 60-61, fig. 21.

Afrogarypus sulcatus rhodesiacus (Beier): Harvey, 1986: 758; Harvey, 1991a: 251.

 

Type locality: Vumba, Zimbabwe.

Distribution: Zimbabwe.

Afrogarypus sulcatus sulcatus (Beier)

 

Distribution: Democratic Republic of Congo (Zaire).

Afrogarypus zonatus (Beier)

Geogarypus (Afrogarypus) zonatus Beier, 1959d: 33-34, fig. 15.

Afrogarypus zonatus (Beier): Harvey, 1986: 758; Harvey, 1991a: 252; Beron, 2002: 33.

 

Type locality: Ht. Ngovi, Itombwe, Uvira, Sud-Kivu, Democratic Republic of Congo (Zaire).

Distribution: Democratic Republic of Congo (Zaire).

Genus Geogarypus Chamberlin

Geogarypus Chamberlin, 1930: 609; Beier, 1932a: 227; Morikawa, 1960: 132; Beier, 1963b: 241-242; Murthy and Ananthakrishnan, 1977: 104-105; Sivaraman, 1980b: 340; Harvey, 1986: 760; Harvey, 1991a: 252; Harvey, 1996a: 259.

Geogarypus (Geogarypus) Chamberlin: Beier, 1932a: 229; Morikawa, 1960: 132; Heurtault, 1970b: 1365; Murthy and Ananthakrishnan, 1977: 105.

 

Type species: Garypus minor L. Koch, 1873, by original designation.

Distribution: AFRICA: widespread.

ASIA: widespread.

AMERICAS: South America; central America.

AUSTRALASIA: Australia; Pacific islands.

EUROPE: Mediterranean region; central Europe.

Geogarypus albus Beier

Geogarypus albus Beier, 1963d: 508-510, fig. 1; Tenorio and Muchmore, 1982: 382; Harvey, 1986: 760; Harvey, 1988b: 329-330, figs 55-58; Harvey, 1991a: 252.

 

Type locality: 8 km N. of Klang, Rantau Panjang, Selangor, Malaysia.

Distribution: Malaysia (mainland).

Geogarypus amazonicus Mahnert

Geogarypus (Geogarypus) amazonicus Mahnert, 1979d: 759-761, figs 85-90; Mahnert, 1985c: 226; Mahnert and Adis, 1986: 213.

Geogarypus amazonicus Mahnert: Adis, 1981: 119, etc.; Mahnert, 1985b: 78; Adis and Mahnert, 1986: 312; Harvey, 1986: 760; Adis, Junk and Penny, 1987: 488; Mahnert, Adis, and Bührnheim, 1986: fig. 6; Harvey, 1991a: 252; Aguiar and Bührnheim, 1994: 74; Morais, Adis, Mahnert and Berti-Filho, 1997: 481; Aguiar and Bührnheim, 1998b: 453; Mahnert and Adis, 2002: 379, fig. 7d; Aguiar and Bührnheim, 2003: 517.

 

Type locality: Manaus, Amazonas, Brazil.

Distribution: Brazil (Amazonas, Pará).

Geogarypus angulatus Chamberlin

Geogarypus angulatus Chamberlin, 1930: 611, figs 2u-v, 3j, 3q; Chamberlin, 1931a: fig. 4b; Harvey, 1986: 760; Harvey, 1991a: 252; Beron, 2002: 34.

Geogarypus (Geogarypus) angulatus Chamberlin: Beier, 1932a: 234-235; Roewer, 1937: 270; Murthy and Ananthakrishnan, 1977: 108.

Geogarypus (Indogarypus) angulatus Chamberlin: Beier, 1957d: 25.

 

Type locality: Ootacamund, Nilgiris, Tamil Nadu, India.

Distribution: India.

Geogarypus asiaticus Murthy and Ananthakrishnan

Geogarypus (Indogarypus) asiaticus Murthy and Ananthakrishnan, 1977: 108-110, figs 36a-b.

Geogarypus asiaticus Murthy and Ananthakrishnan: Harvey, 1986: 760; Harvey, 1991a: 252.

 

Type locality: Chennai (as Madras), Tamil Nadu, India.

Distribution: India.

Geogarypus azerbaidzhanicus Dashdamirov

Geogarypus azerbaidzhanicus Dashdamirov, 1993: 21-23, figs 1a, 1c-d, 1f-g, 2a.

 

Type locality: 5 km W. of Kilyazi, Varafta Mt Ridge, Xizi, Azerbaijan.

Distribution: Azerbaijan.

Geogarypus bucculentus Beier

Geogarypus (Geogarypus) bucculentus Beier, 1955b: 206-207, figs 1-2; Cekalovic, 1984: 16-17.

Geogarypus bucculentus Beier: Beier, 1957c: 454; Harvey, 1986: 760; Harvey, 1987d: 137-140, figs 1-8, 15; Harvey, 1991a: 252.

 

Type locality: Plateau del Yunque, Masatierra, Juan Fernandez Islands, Valparaíso, Chile.

Distribution: Chile (Juan Fernandez Islands).

Geogarypus canariensis (Tullgren)

Garypus canariensis Tullgren, 1900b: 157-159, figs 1-2.

Geogarypus canariensis (Tullgren): Chamberlin, 1930: 609; Beier, 1956e: 306, fig. 3; Beier, 1963b: 243; Beier, 1965c: 379; Beier, 1970a: 45; Beier, 1976a: 24; Estany, 1979a: 222, fig. 1; Pieper, 1981: 3; Bacallado, 1984: 110; Harvey, 1986: 760; Callaini, 1988b: 50-53, figs 5a-g; Harvey, 1991a: 253; Pieper, 1991: 218; Mahnert, 1997: 569.

Geogarypus (Geogarypus) canariensis (Tullgren): Beier, 1932a: 230; Roewer, 1937: 269; Mahnert, 1980a: 259; Mahnert, 1989b: 42.

 

Type locality: Barranco de Ruiz, Tenerife, Islas Canarias, Spain.

Distribution: Morocco; Portugal (Madeira); Spain (Islas Canarias).

Geogarypus ceylonicus Beier

Geogarypus ceylonicus Beier, 1973d: 46, fig. 10; Beier, 1974e: 1008; Harvey, 1986: 760; Harvey, 1991a: 253.

Geogarypus (Indogarypus) ceylonicus Beier: Murthy and Ananthakrishnan, 1977: 110.

 

Type locality: Yakkala, 18 miles NE. of Colombo, Western Province, Sri Lanka.

Distribution: India; Sri Lanka.

Geogarypus connatus Harvey

Geogarypus connatus Harvey, 1986: 772-775, figs 5, 41-53; Harvey, 1991a: 253.

 

Type locality: Horseshoe Bend, Little Desert National Park, Victoria, Australia.

Distribution: Australia (New South Wales, Queensland, Victoria).

Geogarypus continentalis (Redikorzev)

Garypus (Geogarypus) continentalis Redikorzev, 1934a: 432-434, figs 11-12; Roewer, 1936: fig. 66c; Roewer, 1937: 270, fig. 219.

Geogarypus continentalis (Redikorzev): Roewer, 1936: fig. 70n: Redikorzev, 1949: 645; Schawaller, 1985a: 9-10, figs 19-21; Harvey, 1986: 760; Schawaller, 1989: 16; Harvey, 1991a: 253; Dashdamirov and Schawaller, 1993a: 14, fig. 38; Dashdamirov and Schawaller, 1993b: 14; Dashdamirov, 1993: figs 1b, 1e, 2b; Beron, 2002: 34.

Geogarypus irrugatus (Simon): Mahnert, 1977a: 94 (misidentification).

Geogarypus aff. continentalis (Redikorzev): Dashdamirov, 2005: 233-235, figs 41-44.

 

Type locality: Tuyuk Vostochnyy (as Tujuk), Kirghizsky Mountain Ridge, Chüy, Kyrgyzstan.

Distribution: Kazakhstan; Kyrgyzstan; Pakistan?.

Geogarypus cuyabanus (Balzan)

Garypus cuyabanus Balzan, 1887a: no pagination, figs; Balzan, 1890: 441-443, figs 22, 22a; Balzan, 1892: 549.

Geogarypus cuyabanus (Balzan): Chamberlin, 1930: 609; Beier, 1974d: 899; Harvey, 1986: 760; Harvey, 1991a: 253.

Geogarypus (Geogarypus) cuyabanus (Balzan): Beier, 1932a: 231-232; Roewer, 1937: 270.

Not Garypus cuyabanus Balzan: Ellingsen, 1910a: 316 (misidentification; see Geogarypus fiebrigi Beier).

 

Type locality: Mato Grosso (as Mato-Groso), Brazil.

Distribution: Brazil (Mato Grosso, Santa Catarina).

Geogarypus elegans (With)

Garypus personatus Simon, 1901: 79 (junior primary homonym of Garypus personatus Simon, 1900a).

Garypus elegans With, 1906: 107-109, fig. 16, plate 2 figs 2a-b (replacement name for Garypus personatus Simon, 1901).

Geogarypus (Geogarypus) elegans (With): Beier, 1932a: 234; Roewer, 1937: 270.

Geogarypus elegans (With): Harvey, 1986: 760; Harvey, 1991a: 253; Judson, 1997a: 34.

 

Type locality: Kuala Aring, Kelantan, Malaysia.

Distribution: Malaysia (mainland).

Geogarypus exochus Harvey

Geogarypus exochus Harvey, 1986: 769-771, figs 5, 35-37; Harvey, 1991a: 254.

 

Type locality: 90 miles S. of Mackay, Queensland, Australia.

Distribution: Australia (Northern Territory, Queensland).

Geogarypus fiebrigi Beier

Garypus cuyabanus Balzan: Ellingsen, 1910a: 316 (misidentification).

Geogarypus fiebrigi Beier, 1931c: 316; Vachon, 1934i: 414; Harvey, 1986: 760; Harvey, 1991a: 254.

Geogarypus (Geogarypus) fiebrigi Beier: Beier, 1932a: 232, fig. 259; Roewer, 1937: 270.

 

Type locality: San Bernardino, Central, Paraguay.

Distribution: Paraguay.

Geogarypus formosus (Mello-Leitão)

Ideobisium formosum Mello-Leitão, 1937: 269-270, fig. 1; Kury and Nogueira, 1999: 10.

Geogarypus formosus (Mello-Leitão): Mahnert, 1979d: 761; Harvey, 1986: 760; Harvey, 1991a: 254.

 

Type locality: Morro de Caxambú, Petrópolis (as Petropolis), Rio de Janeiro, Brazil.

Distribution: Brazil (Rio de Janeiro).

Geogarypus globulus Sivaraman

Geogarypus (Indogarypus) globulus Sivaraman, 1980b: 340-342, figs 7a-b.

Geogarypus globulus Sivaraman: Harvey, 1986: 760; Harvey, 1991a: 254.

 

Type locality: Pondicherry, Pondicherry, India.

Distribution: India.

† Geogarypus gorskii Henderickx

Geogarypus gorskii Henderickx, 2005: 88-92, figs 1-4.

 

Type locality: Gdansk (as Danzig), Gdansk, Poland [Baltic Amber (Eocene)].

Distribution: Baltic Amber (Eocene).

Geogarypus granulatus Murthy and Ananthakrishnan

Geogarypus (Geogarypus) granulatus Murthy and Ananthakrisnan, 1977: 105-108, figs 35a-b.

Geogarypus granulatus Murthy and Ananthakrishnan: Harvey, 1986: 760; Harvey, 1991a: 254.

 

Type locality: Courtallam, Tamil Nadu, India.

Distribution: India.

Geogarypus heterodentatus Murthy and Ananthakrishnan

Geogarypus (Indogarypus) heterodentatus Murthy and Ananthakrishnan, 1977: 111-112, figs 37a-b.

Geogarypus heterodentatus Murthy and Ananthakrishnan: Sivaraman, 1981: 22; Harvey, 1986: 760; Harvey, 1991a: 254.

 

Type locality: Chennai (as Madras), Tamil Nadu, India.

Distribution: India.

Geogarypus hungaricus (Tömösváry)

Garypus hungaricus Tömösváry, 1882b: 212-213, plate 2 figs 19-21; Daday, 1918: 2.

Geogarypus (Geogarypus) (?) hungaricus (Tömösváry): Beier, 1932a: 235; Roewer, 1937: 270.

Geogarypus (?) hungaricus (Tömösváry): Harvey, 1986: 760.

Geogarypus hungaricus (Tömösváry): Harvey, 1991a: 254.

 

Type locality: Sátoraljaújhely (as Sátoralja-Ujhely), Borsod-Abauj-Zemplén, Hungary.

Distribution: Hungary.

Geogarypus incertus Caporiacco

Geogarypus incertus Caporiacco, 1947b: 21; Caporiacco, 1948a: 617, fig. 10; Harvey, 1986: 760; Harvey, 1991a: 254.

 

Type locality: Guyana.

Distribution: Guyana.

Geogarypus irrugatus (Simon)

Garypus irrugatus Simon, 1899a: 122.

Geogarypus (Geogarypus) irrugatus (Simon): Beier, 1932a: 233 (in part); Roewer, 1937: 270, fig. 221 (in part).

Geogarypus irrugatus (Simon): Roewer, 1936: figs 40a-b, 42c-d (in part); Beier, 1976e: 99-100; Harvey, 1986: 760 (in part); Harvey, 1991a: 255 (in part); Beron, 2002: 34 (in part).

Not Garypus irrugatus Simon: With, 1906: 28-29, 104-107, figs 8a-b, plate 1 figs 6a-d, plate 2 figs 1a-d (misidentification; see Geogarypus longidigitatus (Rainbow)).

Not Geogarypus irrugatus (Simon): Chamberlin, 1930: 611 (misidentification; see Geogarypus longidigitatus (Rainbow)); Chamberlin, 1931a: 114, figs 9m, 15o, 16g, 17z, 32k-l, 37w (misidentification; see Geogarypus longidigitatus (Rainbow)); Mahnert, 1977a: 94 (misidentification, see Geogarypus continentalis (Redikorzev)).

 

Type locality: Sumatra, Indonesia.

Distribution: Bhutan; Indonesia (Sumatra).

Geogarypus longidigitatus (Rainbow)

Chelifer longidigitatus Rainbow, 1897: 108-109, fig. 2.

Garypus longidigitatus (Rainbow): Pocock, 1898: 323; With, 1906: fig. 3; With, 1907: 66-68, fig. 20.

Garypus personatus Simon, 1900a: 518-519.

Garypus javanus Tullgren, 1905: 43-44; Tullgren, 1907a: 66; Ellingsen, 1910a: 388; Ellingsen, 1912a: 122.

Garypus irrugatus Simon: With, 1906: 28-29, 104-107, figs 8a-b, plate 1 figs 6a-d, plate 2 figs 1a-d (misidentification).

Geogarypus javanus (Tullgren): Chamberlin, 1930: 609; Beier, 1953g: 82; Weidner, 1959: 115; Beier, 1982: 43; Harvey, 1986: 760; Harvey, 1988b: 327-329, figs 50-54; Harvey, 1991a: 255; Schawaller, 1994: 733; Schawaller, 1995b: 1050.

Geogarypus personatus (Simon): Chamberlin, 1930: 609; Harvey, 1986: 760; Harvey, 1991a: 259; Judson, 1997a: 34.

Geogarypus longidigitatus (Rainbow): Chamberlin, 1930: 609; Roewer, 1936: figs 22d, 70m; Harvey, 1986: 760; Harvey, 1991a: 256; Harvey, 2000a: 377-383, figs 1-3; Cook, 2003: 2.

Geogarypus irrugatus (Simon): Chamberlin, 1930: 611 (misidentification); Chamberlin, 1931a: 114, figs 9m, 15o, 16g, 17z, 32k-l, 37w (misidentification).

Geogarypus formosanus Beier, 1931c: 315-316, fig. 10 (synonymised by Harvey, 1988b: 327).

Geogarypus (Geogarypus) formosanus Beier: Beier, 1932a: 232-233, fig. 260; Roewer, 1937: 270.

Geogarypus (Geogarypus) longidigitatus (Rainbow): Beier, 1932a: 233; Chamberlin, 1934b: 6; Roewer, 1937: 270.

Geogarypus (Geogarypus) javanus (Tullgren): Beier, 1932a: 233-234, fig. 261; Roewer, 1937: 270; Morikawa, 1963: 6-7, figs c-e.

Geogarypus (Geogarypus) (?) personatus (Simon): Beier, 1932a: 236.

Geogarypus (Geogarypus) personatus (Simon): Chamberlin, 1934b: 6.

Geogarypus (Geogarypus) marquesianus Chamberlin, 1939b: 208-210, figs 1a-g; Tenorio and Muchmore, 1982: 383.

Geogarypus audyi Beier, 1952a: 103-105, fig. 6 (synonymised by Harvey, 1988b: 327).

Geogarypus (Geogarypus) micronesiensis Morikawa, 1952a: 245, figs 4, 5c; Beier, 1957d: 24; Morikawa, 1960: 132-133, plate 3 fig. 2, plate 7 fig. 15; Morikawa, 1962: 419.

Geogarypus (Geogarypus) javanus formosanus Beier: Beier, 1957d: 21-24, figs 9a-b, 10a-d, 11a-d.

Geogarypus (Geogarypus) javanus javanus (Tullgren): Beier, 1957d: 25; Beier, 1965g: 764; Hammen, 1983: 13, 29.

Geogarypus (Geogarypus) javanus audyi Beier: Beier, 1957d: 25; Harvey, 1986: 760.

Geogarypus (Geogarypus) elegans audyi Beier: Beier, 1963d: 507-508.

Geogarypus javanus javanus (Tullgren): Beier, 1966d: 140; Beier, 1970g: 318.

Geogarypus (Geogarypus) javanus takensis Beier, 1967g: 352, fig. 12 (synonymised by Harvey, 1988b: 328).

Geogarypus javanus formosanus Beier: Harvey, 1986: 760.

Geogarypus javanus takensis Beier: Harvey, 1986: 760.

Geogarypus marquesianus Chamberlin: Harvey, 1986: 760; Harvey, 1991a: 256; Benton and Lehtinen, 1995a: 11; Benton and Lehtinen, 1995b: 267.

Geogarypus micronesiensis Morikawa: Roth and Brown, 1976: 128; Harvey, 1986: 760; Harvey, 1991a: 256; Sato, 1994b: 80.

 

Type locality: of Chelifer longidigitatus: Funafuti, Tuvalu.

of Garypus personatus: Kaala Mountains, Oahu Island, Hawaii, U.S.A.

of Garypus javanus: Bogor (as Buitenzorg), Jawa Barat, Java, Indonesia.

of Geogarypus formosanus: Kao-hsiung (as Takao), Kao-hsiung, Taiwan.

of Geogarypus (Geogarypus) marquesianus: Vaipee Valley, Putatauua, Uahuka, Îles Marquises, French Polynesia.

of Geogarypus audyi: Kuala Lumpur, Selangor, Malaysia.

of Geogarypus (Geogarypus) micronesiensis: Minami-tori-shima (as Marcus Island), Japan.

of Geogarypus (Geogarypus) javanus takensis: road to Tak near Thoen, 65 km from Tak, Thailand.

Distribution: Australia (Queensland); Chile (Isla de Pascua); China (Fukien); Cook Islands; Federated States of Micronesia; Fiji; French Polynesia (Archipel des Tuamotu; Îles Marquises, Îles de la Société); Indonesia (Java, Krakatau Islands, Maluku, Sulawesi, Sumba, Timor, West Papua); Japan (Marcus Island); Malaysia; Marshall Islands; Northern Mariana Islands; Papua New Guinea; Pitcairn Islands; Philippines; Samoa; Singapore; Solomon Islands; Taiwan; Thailand; Tonga; Tuvalu; U.S.A. (Hawaii).

† Geogarypus macrodactylus Beier

Geogarypus macrodactylus Beier, 1937d: 305, fig. 4; Roewer, 1940: 331; Beier, 1955g: 51; Schawaller, 1978: 3; Harvey, 1986: 760; Harvey, 1991a: 256; Selden, 1993b: 304.

 

Type locality: Baltic Amber (Eocene).

Distribution: Baltic Amber (Eocene).

Geogarypus maculatus (With)

Garypus maculatus With, 1907: 168-170, figs 21-25.

Geogarypus maculatus (With): Chamberlin, 1930: 609; Harvey, 1986: 760; Harvey, 1991a: 256; Judson, 1997a: 25.

Geogarypus (Geogarypus) maculatus (With): Beier, 1932a: 232; Roewer, 1937: 270.

 

Type locality: Balthazar Estate (as Balthazar), Saint Andrew, Grenada.

Distribution: Grenada.

† Geogarypus major Beier

Geogarypus (?) major Beier, 1937d: 306, fig. 5; Roewer, 1940: 331; Schawaller, 1978: 3; Selden, 1993b: 304.

Geogarypus major Beier: Harvey, 1986: 760; Harvey, 1991a: 256.

 

Type locality: Baltic Amber (Eocene).

Distribution: Baltic Amber (Eocene).

Geogarypus maroccanus Beier

Geogarypus (Geogarypus) maroccanus Beier, 1961b: 31-32, fig. 3.

Geogarypus maroccanus Beier: Harvey, 1986: 760; Callaini, 1988b: 47-50, figs 4a-g; Harvey, 1991a: 256.

 

Type locality: S. of Chechaouen (as Xauen), Chaouen, Morocco.

Distribution: Morocco.

Geogarypus minor (L. Koch)

Garypus minor L. Koch, 1873: 38-40; Simon, 1878a: 152; Simon, 1879a: 46-47, plate 17 fig. 3; Becker, 1880b: cxli; Canestrini, 1884: no. 8, figs 1-4; Daday, 1888: 126, 182; Simon, 1898c: 3; Simon, 1898d: 21; Simon, 1900b: 593; Planet, 1905: unnumbered fig. on p. 284; Tullgren, 1907c: 4-7, fig. 2; Ellingsen, 1908c: 670; Ellingsen, 1909a: 207, 209; Ellingsen, 1910a: 388; Krausse-Heldrungen, 1912: 65; Nonidez, 1917: 27; Navás, 1921: 166; Navás, 1925: 112, fig. 9; Kästner, 1927b: figs 62- 68; Bacelar, 1928: 190; Caporiacco, 1928b: 126; Beier, 1929a: 358; Beier, 1929e: 450; Beier, 1930f: 72; Cloudsley-Thompson, 1968: 139.

Garypus meridionalis Canestrini, 1885: no. 7, figs 1-5 (synonymised by Beier, 1963b: 242).

Cheiridium tetrophthalmum Daday, 1889c: 27-28, figs 8-10, 14; Daday, 1918: 2 (synonymised by Beier, 1932a: 230).

Garypus lusitanus Navás, 1923: 32-33, fig. 12 (synonymised by Navás, 1925: 112).

Geogarypus minor (L. Koch): Chamberlin, 1930: 610; Chamberlin, 1931a: 181; Beier, 1932g: figs 232, 238-245; Roewer, 1937: figs 162-163, 166-176, 178; Beier, 1939f: 196-197; Vachon, 1940g: 144; Beier, 1948a: 190; Caporiacco, 1948b: 34; Beier, 1949: 9; Beier, 1953a: 301; Beier, 1955n: 107; Beier, 1957b: 147; Beier, 1959f: 128; Beier, 1961b: 30; Beier, 1963b: 242; Beier, 1963i: 285; Beier, 1965f: 92; Beier, 1966f: 345; Beier, 1967f: 308; Lazzeroni, 1969a: 335; Lazzeroni, 1969c: 239; Lazzeroni, 1970a: 207; Lagar, 1972a: 20; Gardini, 1975: 8-9; Mahnert, 1975c: 195; Mahnert, 1977d: 97, fig. 4; Callaini, 1979b: 138; Callaini, 1983c: 298-299; Harvey, 1986: 760, fig. 4; Callaini, 1988b: 46; El-Hennawy, 1988b: 10; Harvey, 1991a: 257; Pieper, 1991: 218, 219-220; Gardini, 1995a: 6; Beron, 1996: 24; Judson, 1997a: 27; Mahnert, 1997: 569; Petrov, 1997: 263; García Carrillo, 1998: 12; Gama, Sousa, Ferreira and Barrocas, 2000: 91; Gardini, 2000: 108-109; Kovoor and Muñoz-Cuevas, 2000: 45, 65; Petrov, 2001: 70; Mahnert, 2002: 778; Delfosse, 2003: 37; Mahnert, 2004: 464; Ozimec, 2004: 386; Zaragoza, 2007a: 48.

Geogarypus (Geogarypus) minor (L. Koch): Beier, 1932a: 230, figs 253, 255; Roewer, 1937: 269, fig. 222a; Beier, 1952e: 4.

 

Type locality: of Garypus minor: Corse, France.

of Garypus meridionalis: Rome, Lazio, Italy.

of Cheiridium tetrophthalmum: Vadé, Viana do Castelo, Portugal.

of Garypus lusitanus: São Martinho de Anta (as San Martinho d'Anta), Sabrosa, Vila Real, Portugal.

Distribution: Albania; Algeria; Austria; Bulgaria; Croatia; France (mainland, Corsica); Greece (mainland); Italy (mainland, Sardinia, Sicily); Malta; Morocco; Portugal (Madeira); Spain (mainland, Islas Canarias); Sudan; Turkey.

Geogarypus minutus (Tullgren)

Garypus minutus Tullgren, 1907a: 65-66, fig. 21; Tullgren, 1907b: 229; Ellingsen, 1912b: 87, 108-110; Godfrey, 1927: 18.

Geogarypus minutus (Tullgren): Chamberlin, 1930: 609; Weidner, 1959: 115; Beier, 1964k: 60; Harvey, 1986: 760; Harvey, 1991a: 257.

Geogarypus (Geogarypus) minutus (Tullgren): Beier, 1932a: 231, fig. 257; Roewer, 1937: 269; Beier, 1958a: 171.

 

Type locality: Port Elizabeth, Eastern Cape, South Africa.

Distribution: South Africa.

Geogarypus mirei Heurtault

Geogarypus (?) mirei Heurtault, 1970b: 1361-1365, figs 1-7.

Geogarypus mirei Heurtault: Mahnert, 1978d: 879; Harvey, 1986: 760; Harvey, 1991a: 257.

 

Type locality: Kaortchi, Borkou-Ennedi-Tibesti, Chad.

Distribution: Chad.

Geogarypus nepalensis Beier

Geogarypus nepalensis Beier, 1974b: 268-269, fig. 5; Harvey, 1986: 760; Schawaller, 1987c: 210; Harvey, 1991a: 257; Schawaller, 1991b: 779-780, fig. 1; Beron, 2002: 34.

 

Type locality: between Pina and Rarasee, Nepal.

Distribution: Nepal.

Geogarypus nigrimanus (Simon)

Garypus nigrimanus Simon, 1879a: 47, 312; Simon, 1898c: 3; Simon, 1898d: 21; Simon, 1899d: 86; Simon, 1900b: 592; Berland, 1929: 16.

Geogarypus nigrimanus (Simon): Chamberlin, 1930: 610, figs 1m-n; Chamberlin, 1931a: 227, figs 17x-y, 19k, 44a-b, 62; Beier, 1932g: fig. 258; Vachon, 1936a: 78; Vachon, 1937f: 128; Beier, 1955n: 107; Beier, 1959f: 129; Beier, 1961b: 31; Beier, 1963a: 259; Beier, 1963b: 243, fig. 247; Beier, 1963h: 155; Beier, 1963i: 285; Beier, 1966e: 164; Beier, 1966f: 345; Beier, 1967f: 308; Lazzeroni, 1969a: 335; Lazzeroni, 1969c: 239; Lazzeroni, 1970a: 207; Weygoldt, 1970a: 242, fig. 5; Weygoldt, 1971: 10-12, figs 10-12; Mahnert, 1975c: 195; Orghidan, Dumitresco and Orghidan, 1975: 30-31 (as Geogaripus [sic] nigrimanus); Roth and Brown, 1976: 128; Estany, 1977b: 30-31; Callaini, 1979b: 138-139; Leclerc and Heurtault, 1979: 245; Callaini, 1982a: 17, 20-21, fig. 3b; Pieper, 1981: 3; Callaini, 1982b: 449; Callaini, 1983a: 151-152; Callaini, 1983c: 299-300; Callaini, 1983e: 224; Callaini and Dallai, 1984: 336-341, fig. 5; Harvey, 1986: 760; Callaini, 1988b: 47; Callaini, 1989: 145; Callaini and Dallai, 1989: 90, fig. 13; Dallai and Callaini, 1990: 37-43, figs 1-16; Weygoldt, 1990: fig. 7A5; Harvey, 1991a: 258; Pieper, 1991: 218; Mahnert, 1993a: 357; Gardini, 1995a: 6; Gardini, Sabella and Saccone, 1997: 223; Gardini, 2000: 109; Kovoor and Muñoz-Cuevas, 2000: 45, 65; Delfosse, 2003: 37-38; Šťáhlavský, Král, Harvey and Haddad, 2006: 278, 279, figs 1d-i, 2a, 2c; Zaragoza, 2007a: 49.

Geogarypus (Geogarypus) nigrimanus (Simon): Beier, 1932a: 229, fig. 254; Roewer, 1937: 269.

 

Type locality: Corse, France.

Distribution: Algeria; France (mainland, Corsica); Greece (mainland, Crete, Dodekanisos); Italy (mainland, Sardinia, Sicily); Malta; Morocco; Portugal (Madeira); Spain (mainland, Islas Baleares); Turkey.

Geogarypus ocellatus Mahnert

Geogarypus (Geogarypus) ocellatus Mahnert, 1978d: 874-879, 887, figs 14-20.

Geogarypus ocellatus Mahnert: Harvey, 1986: 760; Harvey, 1991a: 258; Harvey, 2000a: 382; Harvey, 2001b: 691.

 

Type locality: Vallée de Mai, Praslin Island, Seychelles.

Distribution: Seychelles.

Geogarypus olivaceus (Tullgren)

Garypus olivaceus Tullgren, 1907a: 63-65, fig. 20; Tullgren, 1907b: 229.

Garypus senegalensis Balzan: Ellingsen, 1912b: 87, 112-114 (misidentification).

Geogarypus (Geogarypus) olivaceus (Tullgren): Beier, 1932a: 230-231, fig. 256; Roewer, 1937: 269; Beier, 1955l: 301, fig. 20; Beier, 1958a: 170-171.

Geogarypus (Geogarypus) flavus Beier, 1947b: 318-319, fig. 24; Beier, 1958a: 170 (synonymised by Beier, 1964k: 59).

Geogarypus olivaceus (Tullgren): Weidner, 1959: 115; Beier, 1964k: 59; Beier, 1966k: 461, 470; Lawrence, 1967: 89; Harvey, 1986: 760; Harvey, 1991a: 258.

 

Type locality: of Garypus olivaceus: Bothaville, Free State, South Africa.

of Geogarypus (Geogarypus) flavus: Keurbooms River, Knysna, Western Cape, South Africa.

Distribution: South Africa.

Geogarypus palauanus Beier

Geogarypus (Indogarypus) palauanus Beier, 1957d: 25-27, figs 12a-d.

Geogarypus palauanus Beier: Harvey, 1986: 760; Harvey, 1991a: 258.

 

Type locality: Koror, Palau.

Distribution: Palau.

Geogarypus paraguayanus Beier

Geogarypus paraguayanus Beier, 1931c: 316; Harvey, 1986: 760; Harvey, 1991a: 258-259.

Geogarypus (Geogarypus) paraguayanus Beier: Beier, 1932a: 232, fig. 258; Roewer, 1937: 270; Feio, 1945: 4.

 

Type locality: Paraguay.

Distribution: Brazil (Distrito Federal); Paraguay.

Geogarypus pisinnus Harvey

Geogarypus pisinnus Harvey, 1986: 771-772, figs 5, 38-40; Harvey, 1991a: 259.

 

Type locality: Arnhem Highway at South Alligator River, Northern Territory, Australia.

Distribution: Australia (Northern Territory).

Geogarypus pulcher Beier

Geogarypus (Geogarypus) pulcher Beier, 1963f: 192-193, fig. 6; Mahnert, 1974c: 379.

Geogarypus pulcher Beier: Harvey, 1986: 760; Harvey, 1991a: 259.

 

Type locality: Kefar Mallal, HaMerkaz, Israel.

Distribution: Israel.

Geogarypus purcelli (Ellingsen)

Garypus purcelli Ellingsen, 1912b: 110-112.

Geogarypus (Geogarypus) purcelli (Ellingsen): Beier, 1932a: 235; Roewer, 1937: 270; Beier, 1955l: 300, fig. 19.

Geogarypus purcelli (Ellingsen): Beier, 1964k: 59; Beier, 1966k: 461; Harvey, 1986: 760; Harvey, 1991a: 259.

 

Type locality: Beaufort West, Western Cape, South Africa.

Distribution: South Africa.

Geogarypus pustulatus Beier

Geogarypus (Geogarypus) pustulatus Beier, 1959e: 200-201, fig. 13.

Geogarypus pustulatus Beier: Beier, 1962h: 133; Beier, 1964j: 491; Harvey, 1986: 760; Harvey, 1987d: 140-141, figs 9-15; Harvey, 1991a: 259.

 

Type locality: Lago Moreno, San Carlos de Bariloche, Río Negro, Argentina.

Distribution: Argentina; Chile.

Geogarypus quadrimaculatus Mahnert

Geogarypus quadrimaculatus Mahnert, 2007: 285-287, figs 34-38.

 

Type locality: between Hadiboh and Mafarhin, Jo'oh, Suqutra (as Socotra), Yemen.

Distribution: Yemen (Suqutra).

Geogarypus rhantus Harvey

Geogarypus (Geogarypus) rhantus Harvey, 1981a: 279-282, figs 1a-f; Harvey, 1985b: 143.

Geogarypus rhantus Harvey, 1986: 761-764, figs 5-10, 18; Harvey, 1991a: 259.

 

Type locality: Lamond Hill, Iron Range, Queensland, Australia.

Distribution: Australia (Queensland).

Geogarypus robustus Beier

Geogarypus (Geogarypus) robustus Beier, 1947b: 319-320, fig. 25; Beier, 1955l: 301.

Geogarypus robustus (Beier): Harvey, 1986: 760; Harvey, 1991a: 259.

 

Type locality: Fernwood, Western Cape, South Africa.

Distribution: South Africa.

Geogarypus sagittatus Beier

Geogarypus (Indogarypus) sagittatus Beier, 1965g: 764-765, fig. 7; Tenorio and Muchmore, 1982: 383.

Geogarypus sagittatus Beier: Beier, 1982: 43; Harvey, 1986: 760; Harvey, 1991a: 260.

 

Type locality: Ifar, West Papua, Indonesia.

Distribution: Indonesia (West Papua); Papua New Guinea.

Geogarypus shulovi Beier

Geogarypus (Geogarypus) shulovi Beier, 1963f: 193-195, fig. 7; Mahnert, 1974c: 379.

Geogarypus shulovi Beier: Beier, 1965f: 93; Beier, 1967f: 308; Beier, 1971a: 363; Beier, 1973e: 226; Schawaller, 1985a: 10, fig. 22; Harvey, 1986: 760; Harvey, 1991a: 260.

 

Type locality: Daliyya (as Dalia), HaZafon (Northern), Israel.

Distribution: Iran; Israel; Turkey; Turkmenistan.

Geogarypus taylori Harvey

Geogarypus taylori Harvey, 1986: 764-769, figs 5, 11-15, 17, 19-34; Harvey, 1991a: 260; Harvey, 1992c: figs 166-175; Austin, Baker, Colman, Fisher, Harvey, Hirst, Locket and Reay 1996: 122; Šťáhlavský, Král, Harvey and Haddad, 2006: 278, 279-280, figs 2b, 2d.

 

Type locality: Lerderderg Gorge, 9 km NNW. of Bacchus Marsh, Victoria, Australia.

Distribution: Australia (New South Wales, Northern Territory, South Australia, Victoria, Western Australia).

Geogarypus tenuis Chamberlin

Geogarypus tenuis Chamberlin, 1930: 611-612; Chamberlin, 1931a: fig. 29i; Vachon, 1934i: 414; Beier, 1973d: 45; Harvey, 1986: 760; Harvey, 1991a: 260; Judson, 1997a: 43

Geogarypus (Geogarypus) tenuis Chamberlin: Beier, 1932a: 234; Roewer, 1937: 270.

 

Type locality: Sri Lanka? ('presumably from Ceylon').

Distribution: Sri Lanka.

Geogarypus triangularis (Ellingsen)

Garypus minutus triangularis Ellingsen, 1912b: 110.

Geogarypus (Geogarypus) triangularis (Ellingsen): Beier, 1932a: 231; Roewer, 1937: 269.

Geogarypus triangularis (Ellingsen): Beier, 1964k: 59-60, fig. 20; Harvey, 1986: 760; Harvey, 1991a: 260.

 

Type locality: Izeli, near King William's Town, Eastern Cape, South Africa; Bushman's Rock, Blythswood, Butterworth, Eastern Cape, South Africa; and Naval Hill, Bloemfontein, Free State, South Africa.

Distribution: South Africa.

Genus Indogarypus Beier

Geogarypus (Indogarypus) Beier, 1957d: 25; Murthy and Ananthakrishnan, 1977: 108.

Indogarypus Beier: Harvey, 1986: 759; Harvey, 1991a: 260.

 

Type species: Garypus indicus Beier, 1930e, by original designation.

Distribution: ASIA: India; Sri Lanka.

Indogarypus indicus (Beier)

Garypus indicus Beier, 1930e: 290-291, figs 3, 4a-b.

Geogarypus (Geogarypus) indicus (Beier): Beier, 1932a: 235, fig. 262; Roewer, 1937: 270.

Geogarypus (Indogarypus) indicus (Beier): Beier, 1957d: 25; Murthy and Ananthakrishnan, 1977: 108.

Geogarypus indicus (Beier): Beier, 1973d: 46; Beier, 1974e: 1007-1008.

Indogarypus indicus (Beier): Harvey, 1986: 759, fig. 3; Harvey, 1991a: 260.

 

Type locality: Travancore, Kerala, India.

Distribution: India; Sri Lanka.