Pseudoscorpions of the World

Last updated: 19 January 2009

Version 1.2, 2009

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Family ATEMNIDAE Chamberlin

Pessigini Navás, 1925: 109.

Atemnidae Chamberlin, 1931a: 243-244; Beier, 1932b: 548; Beier, 1932e: 20; Beier, 1932g: 186; Roewer, 1937: 279; Hoff, 1956c: 4; Morikawa, 1960: 136; Beier, 1963b: 245; Murthy and Ananthakrishnan, 1977: 124; Muchmore, 1982a: 100; Harvey, 1985b: 128; Harvey, 1991a: 451; Harvey, 1992c: 1430-1431.

Subfamily ATEMNINAE Chamberlin

Atemninae Chamberlin: Beier, 1932b: 549; Beier, 1932e: 20-21; Roewer, 1937: 278; Morikawa, 1960: 136-137; Murthy and Ananthakrishnan, 1977: 124.

Genus Anatemnus Beier

Anatemnus Beier, 1932b: 578; Beier, 1932e: 48; Murthy and Ananthakrishnan, 1977: 129; Harvey, 1991a: 451.

 

Type species: Chelifer javanus Thorell, 1883, by original designation.

Distribution: AFRICA: Madagascar; Seychelles.

AMERICAS: Ecuador; Guyana.

ASIA: widespread.

Anatemnus angustus Redikorzev

Anatemnus angustus Redikorzev, 1938: 96-97, figs 24, 25c; Roewer, 1940: 346; Beier, 1951a: 80-82, fig. 21; Beier, 1955e: 43; Beier, 1967g: 353; Beier, 1976e: 100; Harvey, 1991a: 451; Jędryczkowski, 1998: 186; Beron, 2002: 36; Klausen, 2005: 642-643.

 

Type locality: Djiring, Lâm Dong, Vietnam.

Distribution: Bhutan; India; Malaysia; Vietnam.

Anatemnus elongatus (Ellingsen)

Chelifer (Atemnus) elongatus Ellingsen, 1902: 149-151.

Chelifer elongatus Ellingsen: With, 1908a: 324-326, text-fig. 84, figs 37a-g.

Anatemnus elongatus (Ellingsen): Beier, 1932b: 584; Beier, 1932e: 54; Roewer, 1937: 284; Harvey, 1991a: 451; Klausen, 2005: 643.

 

Type locality: Guayaquil, Guayas, Ecuador.

Distribution: Ecuador; Guyana.

Anatemnus javanus (Thorell)

Chelifer javanus Thorell, 1883: 37-40; With, 1905: 137, plate 8 fig. 1a; With, 1906: 184-185; With, 1907: 59-62, figs 13-14; Ellingsen, 1911b: 141; Ellingsen, 1913b: 127; Ellingsen, 1914: 3Kästner, 1927b: fig. 6.

Chelifer (Lamprochernes) javanus Thorell: Simon, 1901: 79.

Anatemnus javanus (Thorell): Beier, 1932b: 579-580; Beier, 1932e: 49, fig. 58; Roewer, 1937: 283, fig. 201b; Harvey, 1991a: 451-452; Klausen, 2005: 643, fig. 24.

Not Chelifer javanus Thorell: Pocock, 1900: 156 (misidentification; see Paratemnoides pococki (With)).

Not Chelifer (Atemnus) javanus Thorell: Ellingsen, 1910a: 358 (misidentification; see Anatemnus novaguineensis (With)).

 

Type locality: Cibodas (as Tcibodas), Jawa Barat, Java, Indonesia.

Distribution: India; Indonesia (Java); Malaysia; Myanmar; Papua New Guinea; Sri Lanka.

Anatemnus luzonicus Beier

Anatemnus luzonicus Beier, 1932b: 582, fig. 13; Beier, 1932e: 51-52, fig. 62; Roewer, 1937: 283; Harvey, 1991a: 452.

 

Type locality: Mt Data, Bontoc, Mountain, Philippines.

Distribution: Philippines.

Anatemnus madecassus Beier

Chelifer rotundus With: Ellingsen, 1910a: 359 (misidentification); Ellingsen, 1912b: 80 (misidentification).

Anatemnus madecassus Beier, 1932b: 581; Beier, 1932e: 50-51, fig. 60; Roewer, 1937: 283; Millot, 1948a: 142; Legendre, 1972: 446; Harvey, 1991a: 452.

 

Type locality: Andrangoloaka (as Andrangoloaba), Antananarivo, Madagascar.

Distribution: Madagascar.

Anatemnus megasoma (Daday)

Chelifer megasoma Daday, 1897: 476-477, figs 16-17; With, 1906: 171.

Haplochernes megasoma (Daday): Beier, 1932e: 109; Chamberlin, 1934b: 8; Roewer, 1937: 295.

Anatemnus megasoma (Thorell): Beier, 1965g: 769-771, fig. 12; Beier, 1967c: 322; Beier, 1982: 43; Harvey, 1991a: 452.

 

Type locality: Madang (as Friedrich-Wilhelmshafen), Madang, Papua New Guinea.

Distribution: Indonesia (West Papua); Papua New Guinea.

Anatemnus nilgiricus Beier

Atemnus indicus (With): Beier, 1930e: 297-298 (misidentification).

Anatemnus nilgiricus Beier, 1932b: 582-583; Beier, 1932e: 52, fig. 63; Roewer, 1937: 283; Beier, 1967g: 353; Beier, 1973d: 47; Beier, 1974e: 1009; Murthy and Ananthakrishnan, 1977: 129; Harvey, 1991a: 452.

 

Type locality: Nilgiris, Tamil Nadu, India.

Distribution: India; Sri Lanka.

Anatemnus novaguineensis (With)

Chelifer nova-guineensis With, 1908b: 19-22, figs 5-6, plate 2 figs 9-12.

Chelifer (Atemnus) javanus Thorell: Ellingsen, 1910a: 358 (misidentification).

Paratemnus insularis Beier, 1932b: 568; Beier, 1932e: 38, fig. 41; Roewer, 1937: 283 (junior secondary homonym of Paratemnoides insularis (Banks)).

Anatemnus nova-guineensis (With): Beier, 1932b: 584; Beier, 1932e: 53-54; Chamberlin, 1934b: 8; Roewer, 1937: 283.

Paratemnus histrionicus Chamberlin, 1934b: 8; Beier, 1964e: 405-406, fig. 2 (replacement name for Paratemnus insularis Beier; synonymised by Beier, 1965g: 769).

Anatemnus novaguineensis (With): Beier, 1965g: 768-769, fig. 11; Beier, 1966d: 140; Beier, 1966g: 371; Beier, 1970g: 320; Beier, 1982: 43; Harvey, 1991a: 452-453; Judson, 1997a: 30; Klausen, 2005: 643, fig. 23.

 

Type locality: of Chelifer novaguineensis: Blanche Bay, East New Britain, Papua New Guinea.

of Paratemnus insularis: Bismarck Archipelago, Papua New Guinea.

Distribution: Indonesia (West Papua); New Caledonia; Papua New Guinea; Solomon Islands.

Anatemnus orites (Thorell)

Chelifer orites Thorell, 1889: 597-599, plate 5 fig. 7; With, 1906: 188; Tullgren, 1907a: 55-56; Ellingsen, 1911b: 142; Ellingsen, 1914: 4.

Anatemnus orites (Thorell): Beier, 1932b: 580; Beier, 1932e: 50, fig. 59; Roewer, 1937: 283; Beier, 1948b: 461; Weidner, 1959: 114; Harvey, 1991a: 453; Schawaller, 1994: 743-744, figs 38-43; Schawaller, 1995b: 1057; Klausen, 2005: 643.

Not Chelifer orites Thorell: Ellingsen, 1901b: 208-209 (misidentification; see Anatemnus vermiformis (With)); Ellingsen, 1911a: 36 (misidentification; Catatemnus birmanicus (Thorell)).

 

Type locality: Mt Mooleyit, Myanmar.

Distribution: China (Yunnan); India; Indonesia (Sumatra); Malaysia (mainland); Myanmar; Philippines; Sri Lanka; Thailand.

Anatemnus orites major Beier

Anatemnus orites major Beier, 1963d: 510; Harvey, 1991a: 453.

 

Type locality: Rantau Panjang, 5 miles N. of Klang, Selangor, Malaysia.

Distribution: Malaysia (mainland).

Anatemnus orites orites (Thorell)

 

Distribution: China (Yunnan); India; Indonesia (Sumatra); Myanmar; Philippines; Sri Lanka; Thailand.

Anatemnus oswaldi (Tullgren)

Chelifer oswaldi Tullgren, 1907a: 53-55, figs 14a-c (as Chelifer o'swaldi); Ellingsen, 1912b: 80 (as Chelifer o'swaldi).

Atemnus oswaldi (Tullgren): Chamberlin, 1931a: figs 15w, 44j, 44p (as Atemnus o'swaldi); Beier, 1932g: fig. 167 (as Atemnus o'swaldi); Savory, 1935: fig. 63iv; Savory, 1964a: fig. 33d; Savory, 1964a: fig. 93d.

Anatemnus oswaldi (Tullgren): Beier, 1932b: 583 (as Anatemnus o'swaldi); Beier, 1932e: 52-53, fig. 64 (as Anatemnus o'swaldi); Roewer, 1937: 283 (as Anatemnus o'swaldi); Millot, 1948a: 142; Weidner, 1959: 114 (as Anatemnus o'swaldi); Legendre, 1972: 446; Harvey, 1991a: 453.

 

Type locality: Nosy Be (as Nossibé), Antsiranana, Madagascar.

Distribution: Madagascar.

Anatemnus pugilatorius Beier

Anatemnus pugilatorius Beier, 1965g: 768, fig. 10; Harvey, 1991a: 453.

 

Type locality: Finschhafen, Morobe, Papua New Guinea.

Distribution: Papua New Guinea.

Anatemnus rotundus (With)

Chelifer rotundus With, 1906: 196-199, fig. 24, plate 4 figs 11a-c.

Anatemnus rotundus (With): Beier, 1932b: 580-581; Beier, 1932e: 50; Roewer, 1937: 283; Harvey, 1991a: 453.

Not Chelifer rotundus With: Ellingsen, 1910a: 359 (misidentification; see Anatemnus madecassus); Ellingsen, 1912b: 80 (misidentification; see Anatemnus madecassus).

Not Atemnus rotundus (With): Beier, 1930a: 207-208, fig. 5 (misidentification; see Paratemnoides magnificus).

 

Type locality: Little Nicobar (as Nicobar Minor), Nicobar Islands, Andaman and Nicobar Islands, India.

Distribution: India (Nicobar Islands).

Anatemnus seychellesensis Beier

Anatemnus seychellesensis Beier, 1940a: 175-176, fig. 4; Harvey, 1991a: 454; Judson, 1997a: 40; Harvey, 2001b: 691.

 

Type locality: Silhouette Island, Seychelles.

Distribution: Seychelles.

Anatemnus subindicus (Ellingsen)

Chelifer (Atemnus) subindicus Ellingsen, 1910a: 359-360.

Chelifer subindicus Ellingsen: Ellingsen, 1912b: 80.

Anatemnus subindicus (Ellingsen): Beier, 1932b: 581; Beier, 1932e: 51, fig. 61; Roewer, 1937: 283; Millot, 1948a: 142; Legendre, 1972: 446; Harvey, 1991a: 454.

 

Type locality: central Madagascar.

Distribution: Madagascar.

Anatemnus subvermiformis Redikorzev

Anatemnus subvermiformis Redikorzev, 1938: 95-96, figs 23, 25b; Roewer, 1940: 346; Beier, 1951a: 78-80, fig. 20; Harvey, 1991a: 454; Jędryczkowski, 1998: 187; Klausen, 2005: 643.

 

Type locality: Suoi-Tring, Vietnam.

Distribution: Laos; Vietnam.

Anatemnus tonkinensis Beier

Anatemnus angustus tonkinensis Beier, 1943: 76-77, fig. 3.

Anatemnus tonkinensis Beier: Beier, 1951a: 82-83, fig. 22; Beier, 1967g: 353; Harvey, 1991a: 454; Judson, 1997a: 44.

 

Type locality: Ngai-Tio, Tonkin, Vietnam.

Distribution: Laos; Vietnam.

Anatemnus vermiformis (With)

Chelifer orites Thorell: Ellingsen, 1901b: 208-209 (misidentification).

Chelifer vermiformis With, 1906: 188-191, fig. 21, plate 4 figs 8a-b.

Anatemnus vermiformis (With): Beier, 1932b: 585; Beier, 1932e: 54; Roewer, 1937: 283; Beier, 1967g: 353, fig. 13; Harvey, 1991a: 454.

 

Type locality: Nancowry (as Nankovry), Nicobar Islands, Andaman and Nicobar Islands, India.

Distribution: India (mainland and Nicobar Islands); Thailand.

Anatemnus voeltzkowi (Ellingsen)

Chelifer (Atemnus) voeltzkowi Ellingsen, 1908b: 487-488; Ellingsen, 1910a: 362-363.

Chelifer (Atemnus) voeltzkowi elongata Ellingsen, 1908b: 488 (junior primary homonym of Chelifer (Atemnus) elongatus Ellingsen, 1902) (synonymised by Klausen, 2005: 643).

Chelifer voeltzkowi Ellingsen: Ellingsen, 1912b: 81.

Anatemnus voeltzkowi (Ellingsen): Beier, 1932b: 585; Beier, 1932e: 54-55, fig. 66; Roewer, 1937: 284; Millot, 1948a: 142; Legendre, 1972: 446; Harvey, 1991a: 454; Klausen, 2005: 643, fig. 25.

Anatemnus longus Beier, 1932b: 586; Beier, 1932e: 55; Roewer, 1937: 284; Millot, 1948a: 142; Legendre, 1972: 446; Harvey, 1991a: 452.

 

Type locality: of Chelifer (Atemnus) voeltzkowi: south-west Madagascar.

of Chelifer (Atemnus) voeltzkowi elongata: Marovoay, Madagascar.

Distribution: Madagascar.

Genus Atemnus Canestrini

Acis Canestrini, 1883: no. 9 (junior homonym of Acis Billberg, 1820, Acis Lesson, 1830, Acis Dejean, 1835, and Acis Duchassaing and Michelloti, 1860).

Atemnus Canestrini, 1884: no. 1; Beier, 1932b: 574-575; Beier, 1932e: 45; Hoff, 1945c: 1; Beier, 1963b: 246; Murthy and Ananthakrishnan, 1977: 129; Harvey, 1991a: 454-455.

Chelifer (Atemnus) Canestrini: Balzan, 1892: 510; Lessert, 1911: 9-10; Nonidez, 1917: 9.

Pessigus Navás, 1919: 211 (synonymised by Beier, 1932e: 45).

 

Type species: of Acis: Acis brevimanus Canestrini, 1883 (junior synonym of Atemnus politus (Simon, 1878a), by original designation.

of Atemnus: Chelifer politus Simon, 1878a, by subsequent designation of Beier, 1930a: 200.

of Pessigus: Pessigus cabacerolus Navás, 1919 (junior synonym of Atemnus politus (Simon, 1878a)), by present designation.

Distribution: AFRICA: widespread.

AMERICAS: Caribbean.

ASIA: widespread.

EUROPE: widespread.

Atemnus letourneuxi (Simon)

Chelifer letourneuxi Simon, 1881b: 12; Tullgren, 1907c: 3-4, figs 1a-c; Ellingsen, 1912b: 80.

Atemnus letourneuxi (Simon): Simon, 1885c: 49; Pavesi, 1897: 158; Simon, 1897: 158 (not seen); Simon, 1890: 121; Beier, 1932b: 577; Beier, 1932e: 47, fig. 57; Roewer, 1937: 283; Beier, 1965b: 373; El-Hennawy, 1988b: 11; Harvey, 1991a: 455; Mahnert, 1991: 191.

 

Type locality: Mariout, Egypt.

Distribution: Chad; Egypt; Ethiopia; Somalia; Sudan; Tunisia; Yemen.

Atemnus neotropicus Hoff

Atemnus neotropicus Hoff, 1946c: 1-3, figs 1-2; Harvey, 1991a: 455.

 

Type locality: Isla Mona, Puerto Rico.

Distribution: Puerto Rico.

Atemnus politus (Simon)

Chelifer politus Simon, 1878a: 149-150; Simon, 1879a: 35, 312, plate 18 fig. 10; Simon, 1898d: 20; Planet, 1905: 282; Ellingsen, 1909a: 205; Krausse-Heldrungen, 1912: 65; Navás, 1923: 31; Navás, 1925: 106.

Acis brevimanus Canestrini, 1883: no. 10, plate 9 figs 1-4, plate 10 figs 1-4 (synonymised by Ellingsen, 1910a: 358-359).

Atemnus politus (Simon): Canestrini, 1884: no. 1, figs 1-3; Simon, 1885c: 49; Chamberlin, 1931a: 244, figs 14j, 38l, 40v, 41l-n, 42o; Beier, 1932b: 576; Beier, 1932e: 46, figs 54-55; Beier, 1932g: fig. 195; Ionescu, 1936: 4; Roewer, 1937: 283, fig. 227; Vachon, 1938a: figs 26b, 53a; Beier, 1939f: 197; Redikorzev, 1949: 646; Caporiacco, 1951e: 69; Beier, 1953a: 301; Beier, 1955d: 216; Beier, 1955n: 107-108; Beier, 1956e: 303; Beier, 1956i: 8; Beier, 1957b: 147; Beier, 1959a: 270; Beier, 1959c: 407; Beier, 1959f: 129; Schuster, 1959: 44, 65, 66; Beier, 1961b: 32-33; Beier, 1963b: 246-247, fig. 250; Beier, 1963g: 272; Beier, 1963i: 285; Dumitresco and Orghidan, 1964: 625-626, figs 47-48; Orghidan and Dumitresco, 1964a: 328; Orghidan and Dumitresco, 1964b: 192; Beier, 1965f: 93; Kobakhidze, 1965b: 541; Beier, 1966f: 345; Kobakhidze, 1966: 704; Zangheri, 1966: 531; Beier, 1967f: 310; Beier, 1967g: 352-353; Beron, 1968: 104; Beier, 1969b: 193; Dumitresco and Orghidan, 1969: fig. 9; Lazzeroni, 1969a: 335; Lazzeroni, 1969c: 240; Weygoldt, 1969a: 44, 51-52, 62, 75, figs 46, 57; Dumitresco and Orghidan, 1970a: plate 1 fig. 2, plate 2 fig. 6; Lazzeroni, 1970a: 208; Beier, 1971a: 363; Verner, 1971: fig. 50; Weygoldt, 1971: 13, fig. 13; Beier, 1974e: 1008; Ćurčić, 1974a: 26; Gardini, 1976a: 255-256; Mahnert, 1977b: 67-68; Mahnert, 1977d: 97, fig. 5; Murthy and Ananthakrishnan, 1977: 129; Callaini, 1979b: 139; Estany, 1979a: 222, fig. 1; Leclerc and Heurtault, 1979: 246; Callaini, 1982a: 17, 23, fig. 3d; Callaini, 1983c: 300; Callaini, 1983d: 224; Callaini, 1983e: 224; Schawaller, 1983b: 19; Schawaller, 1983c: 371; Bacallado, 1984: 110; Krumpál, 1984b: 63-64; Schawaller, 1985a: 10; Schawaller, 1987c: 212-213; Callaini, 1988b: 55; Schawaller, 1988: 161; Schawaller and Dashdamirov, 1988: 35, figs 56; Callaini, 1989: 145; Schawaller, 1989: 16; Weygoldt, 1990: fig. 8A; Harvey, 1991a: 455-456; Schawaller, 1991b: 782, fig. 1; Dashdamirov and Schawaller, 1992b: 54, fig. 11; Dashdamirov and Schawaller, 1993b: 5; Krumpálova and Krumpál, 1993: 36; Proctor, 1993: 156; Gardini, 1995a: 8; Gardini, 1995b: 48; Schawaller, 1995a: 126; Schawaller, 1995b: 1057; Georgescu and Capuse, 1996a: 111; Petrov, 1997: 266; García Carrillo, 1998: 13; Dashdamirov, 1999: 82; Beron, 2002: 36; Gardini, 2000: 136-137; Delfosse, 2003: 44; Ćurčić, Dimitrijević and Legakis, 2004: 261; Mahnert, 2004: 464; Ozimec, 2004: 388; Petrov, 2004: 159; Dashdamirov, 2005: 247-248; Paul, Donovan and Sudarsanam, 2005: 70; Kárpáthegyi, 2006: 115, fig. 1; Zaragoza, 2007a: 51-52.

Chelifer (Atemnus) politus (Simon): Simon, 1900b: 593; Ellingsen, 1910a: 358-359; Lessert, 1911: 10-11, fig. 9; Kästner, 1928: 5, fig. 13; Beier, 1929a: 342; Beier, 1929b: 78; Beier, 1929e: 445.

Chelifer (Atemnus) ariasi Nonidez, 1917: 9-12, figs 1, 2a-c (synonymised by Beier, 1963b: 247).

Chelifer ariasi Nonidez: Navás, 1918: 133-134.

Chelifer (Atemnus) turkestanicus Redikorzev, 1922a: 257-259, figs 1-2 (synonymised by Krumpál, 1984b: 63).

Pessigus ariasi (Nonidez): Navás, 1919: 211; Navás, 1925: 109.

Pessigus cabacerolus Navás, 1919: 211-212, figs 8a-c; Navás, 1925: 109-110 (synonymised by Beier, 1932e: 46).

Chelifer (Chernes) politus Simon: Berland, 1929: 15.

Atemnus turkestanicus (Redikorzev): Beier, 1932b: 576-577; Beier, 1932e: 47, fig. 56; Roewer, 1937: 283; Redikorzev, 1949: 646; Beier, 1959a: 270; Beier, 1967g: 353; Beier, 1976e: 100; Beron, 2002: 36; Klausen, 2005: 643, fig. 9.

Atemnus ariasi (Nonidez): Beier, 1932b: 577; Beier, 1932e: 48; Roewer, 1937: 283.

Atemnus balcanicus Hadži, 1938: 20-23, figs 22a-d, 23a-b; Ćurčić, 1974a: 26 (synonymised by Beier, 1963b: 247).

Microcreagris zangherii Caporiacco, 1948c: 241-242, fig. 1; Beier, 1963b: 208; Zangheri, 1966: 531; Cerruti, 1968: fig. 7 (synonymised by Gardini, 1976a: 255).

Atemnus politus turkestanicus (Redikorzev): Beier, 1974b: 269; Beier, 1973f: 167.

Not Chelifer politus Simon: Hansen, 1884: 543 (misidentification; see Oratemnus navigator (With)).

 

Type locality: of Chelifer politus: Dhaya (as Daya), Sidi Bel Abbès, Algeria.

of Acis brevimanus: Acireale, Sicily, Italy.

of Chelifer (Atemnus) ariasi: Algeciras, Cádiz, Andalucía, Spain.

of Pessigus cabacerolus: Cabacés, Tarragona, Cataluña, Spain.

of Chelifer (Atemnus) turkestanicus: Tashkent, Tashkent, Uzbekistan.

of Atemnus balcanicus: Treske River, near Manastira Sveti Nikola (as "Klosters Sv. Nikola"), Macedonia.

of Microcreagris zangherii: Pineta di Classe, Emilia-Romagna, Italy.

Distribution: Afghanistan; Algeria; Austria; Azerbaijan; Bhutan; Bulgaria; China (Hupeh); Croatia; France (mainland); Georgia; Greece (mainland, Crete); Hungary; India; Iran; Italy (mainland, Sardinia, Sicily); Kazakhstan; Kyrgyzstan; Libya; Macedonia; Mongolia; Morocco; Nepal; Pakistan; Romania; Slovakia; Spain (mainland, Islas Baleares, Islas Canarias); Switzerland; Syria; Slovakia; Tunisia; Turkey; Turkmenistan; Ukraine; Uzbekistan.

 

Atemnus strinatii Beier

Atemnus strinatii Beier, 1977a: 188-190, fig. 2; Harvey, 1991a: 456; Strinati, 1983: 30.

 

Type locality: Cueva Basilen, Pagbilao, Quezon, Philippines.

Distribution: Philippines.

Atemnus syriacus (Beier)

Catatemnus syriacus Beier, 1955d: 216, figs 4-5; Beier, 1963g: 272; Beier, 1965f: 93.

Atemnus syriacus (Beier): Beier, 1967f: 310; Mahnert, 1977b: 67; Harvey, 1991a: 456; Klausen, 2005: 644.

Atemnus (?) syriacus (Beier): Mahnert, 1974c: 379.

 

Type locality: Al-Lādhiqīyah (as Latakia), Al-Lādhiqīyah, Syria.

Distribution: Greece (mainland); Israel; Syria; Turkey.

Genus Athleticatemnus Beier

Athleticatemnus Beier, 1979c: 101-103; Harvey, 1991a: 456.

 

Type species: Athleticatemnus pugil Beier, 1979c, by original designation.

Distribution: AFRICA: Democratic Republic of Congo (Zaire).

Athleticatemnus pugil Beier

Athleticatemnus pugil Beier, 1979c: 103-104, fig. 1; Harvey, 1991a: 456; Klausen, 2005: 644.

 

Type locality: Luki, Boma, Zaire Central, Democratic Republic of Congo (Zaire).

Distribution: Democratic Republic of Congo (Zaire).

Genus Catatemnus Beier

Catatemnus Beier, 1932b: 593-594; Beier, 1932e: 62-63; Murthy and Ananthakrishnan, 1977: 130; Harvey, 1991a: 457.

 

Type species: Chelifer birmanicus Thorell, 1889, by original designation.

Distribution: AFRICA: widespread.

ASIA: widespread.

Catatemnus birmanicus (Thorell)

Chelifer birmanicus Thorell, 1889: 594-597, plate 5 fig. 6; With, 1906: 176-180, plate 3, fig. 10a, plate 4 figs 4a-f.

Chelifer orites Thorell: Ellingsen, 1911a: 36 (misidentification).

Atemnus birmanicus (Thorell): Chamberlin, 1931a: fig. 18m; Vachon, 1949: fig. 203h.

Catatemnus birmanicus (Thorell): Beier, 1932b: 595; Beier, 1932e: 63-64; Roewer, 1936: figs 22g, 112k, 142; Roewer, 1937: 284; Vachon, 1940f: 3; Weidner, 1959: 113; Murthy and Ananthakrishnan, 1977: 130; Harvey, 1991a: 457; Judson, 1997a: 10; Klausen, 2005: 644, fig. 26.

Not Chelifer birmanicus Thorell: Tullgren, 1905: 39-40, figs 1a-f (misidentification; see Catatemnus thorelli (Balzan)); Tullgren, 1912a: 267 (misidentification; see Paratemnoides assimilis); Tullgren, 1912b: 269 (misidentification; see Paratemnoides assimilis); Dammerman, 1922: 100 (misidentification; see Paratemnoides assimilis); Dammerman, 1929: 114 (not seen) (misidentification; see Paratemnoides assimilis); Bristowe, 1931b: 1390 (misidentification; see Paratemnoides assimilis); Dammerman, 1948: 496 (misidentification; see Paratemnoides assimilis).

 

Type locality: Bhamó, Kachin, Myanmar.

Distribution: Indonesia (Sumatra); Myanmar.

Catatemnus braunsi (Tullgren)

Chelifer braunsi Tullgren, 1907a: 56-58, fig. 16; Tullgren, 1907b: 224; Ellingsen, 1912b: 78.

Catatemnus braunsi (Tullgren): Beier, 1932b: 599-600; Beier, 1932e: 67-68, fig. 82; Roewer, 1937: 284; Beier, 1958a: 173; Weidner, 1959: 113; Beier, 1964k: 72; Harvey, 1991a: 457.

Paratemnus braunsi (Tullgren): Weygoldt, 1970a: 242, figs 13a-b; Schaller, 1979: plate 10.III fig. 12.

Paratemnoides braunsi (Tullgren): Proctor, 1993: 156.

 

Type locality: Algoa Bay, Port Elizabeth, Eastern Cape, South Africa.

Distribution: South Africa; Tanzania.

Catatemnus comorensis (Ellingsen)

Chelifer (Lamprochernes) comorensis Ellingsen, 1910a: 366-367.

Chelifer comorensis Ellingsen: Ellingsen, 1912b: 81.

Catatemnus comorensis (Ellingsen): Beier, 1932b: 597-598; Beier, 1932e: 65-66, fig. 78; Roewer, 1937: 284; Mahnert, 1978f: 125-126; Mahnert, 1983b: 362-364, figs 9-15; Mahnert, 1984b: 60-61; Harvey, 1991a: 458.

 

Type locality: Mayotte.

Distribution: Kenya; Mayotte; Somalia.

Catatemnus concavus (With)

Chelifer concavus With, 1906: 173-176, plate 4 fig. 3a.

Catatemnus concavus (With): Beier, 1932b: 600; Harvey, 1991a: 458.

Catatemnus (?) concavus (With): Beier, 1932e: 68; Roewer, 1937: 284.

 

Type locality: Nancowry (as Nankovry), Nicobar Islands, Andaman and Nicobar Islands, India.

Distribution: India (Nicobar Islands).

Catatemnus exiguus Mahnert

Catatemnus exiguus Mahnert, 1978f: 114-119, figs 108-118; Harvey, 1991a: 458.

 

Type locality: Meya, Tal de Niari, Niari, Congo.

Distribution: Congo.

Catatemnus fravalae Heurtault

Catatemnus fravalae Heurtault, 1983: 18-21, figs 43-47; Harvey, 1991a: 458.

 

Type locality: Lamto, near Toumodi, Lacs, Côte d’Ivoire.

Distribution: Côte d’Ivoire.

Catatemnus granulatus Mahnert

Catatemnus granulatus Mahnert, 1978f: 112-114, figs 103-107; Harvey, 1991a: 458; Klausen, 2005: 644, fig. 27.

 

Type locality: Sibiti, Lékoumou, Congo.

Distribution: Cameroon; Congo; Democratic Republic of Congo (Zaire).

Catatemnus kittenbergeri Caporiacco

Catatemnus kittenbergeri Caporiacco, 1947a: 98, fig. 1; Harvey, 1991a: 458.

 

Type locality: Moshi, Kilimanjaro, Tanzania.

Distribution: Tanzania.

Catatemnus monitor (With)

Chelifer monitor With, 1906: 180-182, plate 4 figs 5a-b; With, 1908b: 15.

Catatemnus monitor (With): Beier, 1932b: 596; Beier, 1932e: 64; Roewer, 1937: 284; Harvey, 1991a: 458.

 

Type locality: 'Sunda Islands', Indonesia.

Distribution: Indonesia.

Catatemnus nicobarensis (With)

Chelifer nicobarensis With, 1906: 182-184, fig. 17, plate 4 figs 6a-b; Ellingsen, 1911a: 37.

Catatemnus nicobarensis (With): Beier, 1932b: 596; Beier, 1932e: 64; Roewer, 1937: 284; Harvey, 1991a: 458.

 

Type locality: Pulu-Mulu, Nicobar Islands, Andaman and Nicobar Islands, India.

Distribution: India (Nicobar Islands); Indonesia (Sumatra).

Catatemnus schlottkei Vachon

Catatemnus schlottkei Vachon, 1937a: 130-132, figs 3-4; Roewer, 1940: 346; Harvey, 1991a: 459.

 

Type locality: Makapau, Victoria, Masvingo, Zimbabwe.

Distribution: Zimbabwe.

Catatemnus thorelli (Balzan)

Chelifer (Lamprochernes) thorelli Balzan, 1891: 519-520, figs 11, 11a.

Chelifer birmanicus Thorell: Tullgren, 1905: 39-40, figs 1a-f (misidentification).

Chelifer thorelli Balzan: With, 1906: 176 (as Chelifer thorellii); Ellingsen, 1909a: 219.

Atemnus thorelli (Balzan): Beier, 1930a: 209.

Catatemnus thorelli (Balzan): Beier, 1932b: 597; Beier, 1932e: 65, figs 76-77; Roewer, 1937: 284; Harvey, 1991a: 459.

 

Type locality: Sumatra, Indonesia.

Distribution: Indonesia (Borneo, Java, Sumatra); Malaysia (Sarawak).

Catatemnus togoensis (Ellingsen)

Chelifer (Lamprochernes) togoensis Ellingsen, 1910a: 368-369.

Chelifer togoensis Ellingsen: Ellingsen, 1910b: 63; Ellingsen, 1910d: 536; Ellingsen, 1912b: 81.

Catatemnus togoensis (Ellingsen): Beier, 1932b: 598-599; Beier, 1932e: 66-67, fig. 80; Roewer, 1937: 284; Vachon, 1941e: 29; Beier, 1947b: 324-325; Vachon, 1952a: 29, fig. 17; Beier, 1954d: 135; Mahnert, 1978f: 119-126, figs 119-130; Mahnert, 1983b: 361; Harvey, 1991a: 459; Klausen, 2005: 644, fig. 28.

Catatemnus congicus Beier, 1932b: 598, fig. 17; Beier, 1932e: 66, fig. 79; Roewer, 1937: 284; Beier, 1947b: 324; Vachon, 1952a: 29; Beier, 1954d: 134; Beier, 1959d: 43 (synonymised by Mahnert, 1978f: 119).

Catatemnus sommerfeldi Beier, 1932b: 599; Beier, 1932e: 67, fig. 81; Roewer, 1937: 284; Vachon, 1941e: 29 (synonymised by Mahnert, 1978f: 119).

Catatemnus similis Vachon, 1938c: 311-313, figs i, m, o-r; Roewer, 1940: 346; Vachon, 1941e: 29 (synonymised by Mahnert, 1978f: 119).

Catatemnus cf. togoensis (Ellingsen): Mahnert, 1978f: 126.

 

Type locality: of Chelifer (Lamprochernes) togoensis: Bismarckbourg, Centrale, Togo.

of Catatemnus congicus: Kikiongo Goma, Maduda, Zaire Central, Democratic Republic of Congo (Zaire).

of Catatemnus sommerfeldi: Yaoundé (as Jaunde), Centre, Cameroon; and Abang Mbang, Est, Cameroon.

of Catatemnus similis: Bingerville, Abidjan, Côte d’Ivoire.

Distribution: Angola; Cameroon; Congo; Côte d’Ivoire; Democratic Republic of Congo (Zaire); Guinea; Kenya; Nigeria; Togo; Uganda.

Genus Cyclatemnus Beier

Cyclatemnus Beier, 1932b: 560; Beier, 1932e: 30-31; Harvey, 1991a: 459.

 

Type species: Cyclatemnus centralis Beier, 1932b, by original designation.

Distribution: AFRICA: widespread.

Cyclatemnus affinis Vachon

Cyclatemnus affinis Vachon, 1938c: 307-308, figs d-f, k; Roewer, 1940: 346; Harvey, 1991a: 459.

 

Type locality: Bingerville, Abidjan, Côte d’Ivoire.

Distribution: Côte d’Ivoire.

Cyclatemnus berlandi Vachon

Cyclatemnus berlandi Vachon, 1938c: 309-311, figs b-c, g, j, n; Roewer, 1940: 346; Harvey, 1991a: 459.

 

Type locality: Dakar, Dakar, Senegal.

Distribution: Senegal.

Cyclatemnus brevidigitatus Mahnert

Cyclatemnus brevidigitatus Mahnert, 1978f: 96, figs 139-144; Harvey, 1991a: 460.

 

Type locality: Lefini reservation, Nanbouli River, Plateaux, Congo.

Distribution: Congo.

Cyclatemnus burgeoni (Beier)

Paratemnus burgeoni Beier, 1932b: 571; Beier, 1932e: 41-42, fig. 47-48; Roewer, 1937: 283; Vachon, 1941e: 30.

Cyclatemnus burgeoni (Beier): Beier, 1944: 184-185, fig. 9; Beier, 1954d: 134; Beier, 1955a: 12; Beier, 1959d: 40; Beier, 1972a: 11; Harvey, 1991a: 460; Klausen, 2005: 644.

 

Type locality: Moto, Uele, Democratic Republic of Congo (Zaire).

Distribution: Cameroon; Democratic Republic of Congo (Zaire); Uganda.

Cyclatemnus centralis Beier

Cyclatemnus centralis Beier, 1932b: 561, fig. 6; Beier, 1932e: 31-32, figs 30-31; Roewer, 1937: 282; Beier, 1967d: 79; Mahnert, 1983b: 375-378, figs 48-57; Harvey, 1991a: 460; Beron, 2002: 36; Klausen, 2005: 644-645, fig. 21.

 

Type locality: south-west Rwanda.

Distribution: Democratic Republic of Congo (Zaire); Kenya; Rwanda.

Cyclatemnus dolosus Beier

Cyclatemnus globosus Beier: Beier, 1958a: 172-173, fig. 7 (misidentification).

Cyclatemnus dolosus Beier, 1964k: 71; Beier, 1966k: 464; Harvey, 1991a: 460; Klausen, 2005: 645; Haddad, Dippenaar-Schoeman and Wesołowska, 2006: 21.

 

Type locality: Town Bush, near Pietermaritzburg, KwaZulu-Natal, South Africa.

Distribution: South Africa; Zimbabwe.

Cyclatemnus equestroides (Ellingsen)

Chelifer (Atemnus) equestroides Ellingsen, 1906: 250-251; Ellingsen, 1910a: 367; Ellingsen, 1912b: 79.

Tamenus (?) equestroides (Ellingsen): Beier, 1932b: 603-604; Beier, 1932e: 72; Roewer, 1937: 285.

Tamenus equestroides (Ellingsen): Harvey, 1991a: 477.

Cyclatemnus equestroides (Ellingsen): Klausen, 2005: 645.

 

Type locality: Rio Cassine, Guinea-Bissau; Ribeira Palma, São Tomé (as Thomé), São Tomé and Príncipe; Roça, Príncipe, São Tomé and Príncipe; Bahia do Oeste, Príncipe, São Tomé and Príncipe; Punta Frailes, Fernando Poo (now Macias Nguema), Equatorial Guinea; and Lake N'Komi (as Fernand Vaz), Ogooué-Maritime, Gabon.

Distribution: Equatorial Guinea; Gabon; Guinea-Bissau; São Tomé and Príncipe.

Cyclatemnus fallax Beier

Cyclatemnus fallax Beier, 1955c: 549-551, fig. 15; Beier, 1967d: 80; Mahnert, 1983b: 378-380, figs 58-62; Harvey, 1991a: 460; Beron, 2002: 36.

 

Type locality: Suam River, Mt Elgon, Uganda.

Distribution: Kenya; Uganda.

Cyclatemnus globosus Beier

Cyclatemnus globosus Beier, 1947b: 323-324, fig. 29; Beier, 1966k: 470 (as Cyclotemnus [sic] globosus); Harvey, 1991a: 460; Klausen, 2005: 645.

Not Cyclatemnus globosus Beier: Beier, 1958a: 172-173, fig. 7 (misidentification; see Cyclatemnus dolosus Beier).

 

Type locality: Kakamas, Orange River, Northern Cape, South Africa.

Distribution: Namibia; South Africa.

Cyclatemnus globosus globosus Beier

 

Distribution: Namibia; South Africa.

Cyclatemnus globosus parvus Beier

Cyclatemnus globosus parvus Beier, 1964k: 70-71, fig. 28; Beier, 1966k: 464; Lawrence, 1967: 89; Harvey, 1991a: 460; Haddad, Dippenaar-Schoeman and Wesołowska, 2006: 21.

 

Type locality: Roodeplaat, Pretoria, Gauteng, South Africa.

Distribution: South Africa.

Cyclatemnus granulatus Beier

Cyclatemnus granulatus Beier, 1932b: 561-562; Beier, 1932e: 32, fig. 32; Roewer, 1937: 282; Vachon, 1941e: 29; Vachon, 1952a: 30; Beier, 1954d: 134; Heurtault, 1983: 18; Harvey, 1991a: 460; Klausen, 2005: 645.

 

Type locality: Bosum, Cameroon.

Distribution: Cameroon; Côte d’Ivoire; Democratic Republic of Congo (Zaire); Guinea.

Cyclatemnus minor Beier

Cyclatemnus minor Beier, 1944: 185-187, fig. 10; Beier, 1967d: 80, fig. 5; Mahnert, 1983b: 375; Harvey, 1991a: 461; Judson, 1997a: 26-27; Beron, 2002: 36; Klausen, 2005: 645.

 

Type locality: Jemme Forest (as Djem-Djem Wald), Ethiopia.

Distribution: Ethiopia; Kenya.

Cyclatemnus robustus Beier

Cyclatemnus robustus Beier, 1959d: 40-41, fig. 19; Harvey, 1991a: 461; Beron, 2002: 36 Klausen, 2005: 645.

 

Type locality: Mwenga, Sud-Kivu, Democratic Republic of Congo (Zaire).

Distribution: Democratic Republic of Congo (Zaire).

Genus Mesatemnus Beier and Turk

Anatemnus (Mesatemnus) Beier and Turk, 1952: 769-770.

Mesatemnus Beier and Turk: Beier, 1963b: 247; Harvey, 1991a: 463.

 

Type species: Anatemnus (Mesatemnus) cyprianus Beier and Turk, 1952, by original designation

Distribution: EUROPE: Cyprus.

Mesatemnus cyprianus (Beier and Turk)

Anatemnus (Mesatemnus) cyprianus Beier and Turk, 1952: 770-771, figs 3-5.

Mesatemnus cyprianus (Beier and Turk): Beier, 1963b: 247; Harvey, 1991a: 463.

 

Type locality: Salamis, Alajuela, Cyprus.

Distribution: Cyprus.

Genus Metatemnus Beier

Metatemnus Beier, 1932b: 606-607; Beier, 1932e: 75; Harvey, 1991a: 463.

 

Type species: Metatemnus philippinus Beier, 1932b, by original designation.

Distribution: ASIA: widespread.

Metatemnus heterodentatus Beier

Metatemnus heterodentatus Beier, 1952a: 106-108, figs 9-10; Harvey, 1991a: 463; Schawaller, 1994: 744-745.

 

Type locality: Mehipit, Borneo.

Distribution: "Borneo"; Thailand.

Metatemnus philippinus Beier

Metatemnus philippinus Beier, 1932b: 607-608; Beier, 1932e: 75-76, figs 93-95; Roewer, 1937: 285, fig. 228; Beier, 1965g: 772; Beier, 1966b: 343; Harvey, 1991a: 464.

Metatemnus major Beier, 1932b: 608; Beier, 1932e: 76, fig. 96; Roewer, 1937: 285 (synonymised by Beier, 1965g: 772).

 

Type locality: of Metatemnus philippinus: Port Banga, Zamboanga del Sur, Mindanao Island, Philippines.

of Metatemnus major: Sabaan, Ilocos Sur, Luzon Island, Philippines.

Distribution: Indonesia (West Papua); Papua New Guinea; Philippines.

Metatemnus superior Muchmore

Metatemnus superior Muchmore, 1972a: 11-14, figs 1-3; Harvey, 1991a: 464; Poinar, Ćurčić and Cokendolpher, 1998: 85.

 

Type locality: Cameron Highlands, Pahang, Malaysia.

Distribution: Malaysia (mainland).

Metatemnus unistriatus (Redikorzev)

Anatemnus unistriatus Redikorzev, 1938: 97-99, fig. 26; Roewer, 1940: 346.

Metatemnus unistriatus (Redikorzev): Beier, 1951a: 86; Harvey, 1991a: 464; Jędryczkowski, 1998: 187.

 

Type locality: Da Lat (as Dalat), Lâm Dong, Vietnam.

Distribution: Vietnam.

Genus Micratemnus Beier

Micratemnus Beier, 1932b: 587; Beier, 1932e: 56; Murthy and Ananthakrishnan, 1977: 130; Harvey, 1991a: 464.

 

Type species: Chelifer (Atemnus) pusillus Ellingsen, 1906, by original designation.

Distribution: AFRICA: widespread.

ASIA: widespread.

Micratemnus anderssoni Beier

Micratemnus anderssoni Beier, 1973d: 49, fig. 13; Harvey, 1991a: 464.

 

Type locality: Deerwood Kuruwita, 6 miles NNW. of Ratnapura, Sabaragamuwa Province, Sri Lanka.

Distribution: Sri Lanka.

Micratemnus ceylonicus Beier

Micratemnus ceylonicus Beier, 1973d: 47-49, fig. 12; Beier, 1974e: 1009-1010, fig. 6; Murthy and Ananthakrishnan, 1977: 130; Harvey, 1991a: 464; Schawaller, 1994: 745.

 

Type locality: Yalakumbura, 5 miles SSW. of Bibile, Uva Province, Sri Lanka.

Distribution: India; Sri Lanka; Thailand.

Micratemnus crassipes Mahnert

Micratemnus sulcatus Beier: Beier, 1967d: 81 (misidentification).

Micratemnus crassipes Mahnert, 1983b: 366-368, figs 22-29; Mahnert, 1984b: 60; Harvey, 1991a: 464; Klausen, 2005: 645.

 

Type locality: Lake Baringo Lodge, Lake Baringo, Rift Valley, Kenya.

Distribution: Kenya; Somalia.

Micratemnus pusillus (Ellingsen)

Chelifer (Atemnus) pusillus Ellingsen, 1906: 248-250.

Chelifer pusillus Ellingsen: Ellingsen, 1912b: 80; Ellingsen, 1913a: 452.

Micratemnus pusillus (Ellingsen): Beier, 1932b: 587-588; Beier, 1932e: 56-57, fig. 68; Roewer, 1937: 284; Beier, 1948b: 452; Harvey, 1991a: 464.

 

Type locality: Vista Alegre, São Tomé, São Tomé and Príncipe.

Distribution: Ghana; São Tomé and Príncipe.

Micratemnus sulcatus Beier

Micratemnus sulcatus Beier, 1944: 187-188, fig. 11; Mahnert, 1983b: 364-366, figs 16-21; Harvey, 1991a: 465; Beron, 2002: 36.

Not Micratemnus sulcatus Beier: Beier, 1967d: 81 (misidentification; see Micratemnus crassipes Mahnert).

 

Type locality: Tanga, Tanga, Tanzania.

Distribution: Kenya; Tanzania.

Genus Oratemnus Beier

Oratemnus Beier, 1932b: 588; Beier, 1932e: 57; Morikawa, 1960: 138; Beier, 1976f: 213; Murthy and Ananthakrishnan, 1977: 126; Sivaraman, 1980c: 349; Harvey, 1991a: 465.

Steiratemnus Beier, 1948c: 527-529 (synonymised by Beier, 1966a: 296).

 

Type species: of Oratemnus: Chelifer (Lamprochernes) articulosus Simon, 1899a, by original designation.

of Steiratemnus: Chelifer punctatus L. Koch, 1885, by original designation.

Distribution: AFRICA: Seychelles.

AMERICAS: Caribbean.

ASIA: widespread.

AUSTRALASIA: Australia.

Oratemnus afghanicus Beier

Oratemnus afghanicus Beier, 1959a: 270-271, fig. 13; Harvey, 1991a: 466.

 

Type locality: Darountah, Jalalabad (as Djelalabad), Afghanistan.

Distribution: Afghanistan.

Oratemnus articulosus (Simon)

Chelifer (Lamprochernes) articulosus Simon, 1899a: 120-121.

Chelifer articulosus Simon: With, 1906: 196; Ellingsen, 1911a: 36.

Oratemnus articulosus (Simon): Beier, 1932b: 589; Beier, 1932e: 58, fig. 69; Roewer, 1937: 284; Harvey, 1991a: 466.

 

Type locality: Sumatra, Indonesia.

Distribution: Indonesia (Sumatra).

Oratemnus boettcheri Beier

Oratemnus boettcheri Beier, 1932b: 592, fig. 15 (as Oratemnus böttcheri); Beier, 1932e: 60-61, fig. 73 (as Oratemnus böttcheri); Roewer, 1937: 284 (as Oratemnus böttcheri); Beier, 1966b: 343 (as Oratemnus böttcheri); Beier, 1967g: 354; Harvey, 1991a: 466; Jędryczkowski, 1998: 187.

 

Type locality: Vinac Island, Philippines.

Distribution: Philippines; Vietnam.

Oratemnus brevidigitatus Beier

Oratemnus brevidigitatus Beier, 1940a: 176-177, fig. 5; Beier, 1948b: 442, 457; Mahnert, 1978d: 880-882, figs 24-29; Harvey, 1991a: 466; Judson, 1997a: 11; Poinar, Ćurčić and Cokendolpher, 1998: 85; Harvey, 2001b: 691.

 

Type locality: Mahé Island, Seychelles.

Distribution: Seychelles.

Oratemnus cavernicola Beier

Oratemnus cavernicola Beier, 1976b: 271-272, fig. 1; Harvey, 1981b: 245; Harvey, 1985b: 128; Harvey, 1991a: 466; Moulds, 2004: 12.

 

Type locality: Jump Up Cave, Gray Range, ca. 48 km N. of Tibooburra, New South Wales, Australia.

Distribution: Australia (New South Wales).

Oratemnus confusus Murthy and Ananthakrishnan

Oratemnus confusus Murthy and Ananthakrishnan, 1977: 127-129, figs 41a-b; Harvey, 1991a: 466.

 

Type locality: Coimbatore, Tamil Nadu, India.

Distribution: India.

Oratemnus curtus (Beier)

Steiratemnus curtus Beier, 1954b: 15-16, fig. 6.

Oratemnus curtus (Beier): Beier, 1966a: 278; Harvey, 1981b: 245; Harvey, 1985b: 128; Harvey, 1991a: 466; Postle, Majer and Bell, 1991: 196; Harvey, 1992c: figs 235-244.

 

Type locality: Pemberton, Western Australia, Australia.

Distribution: Australia (Western Australia).

Oratemnus distinctus (Beier)

Steiratemnus distinctus Beier, 1948c: 531-532, fig. 4.

Oratemnus distinctus (Beier): Beier, 1966a: 278; Beier, 1969a: 187; Richards, 1971: 18; Harvey, 1981b: 245; Harvey, 1985b: 128; Harvey, 1991a: 466; Judson, 1997a: 15.

 

Type locality: New South Wales, Australia.

Distribution: Australia (New South Wales, Western Australia).

Oratemnus indicus (With)

Chelifer indicus With, 1906: 194-196, fig. 23, plate 4 figs 10a-d; Ellingsen, 1914: 3; Roewer, 1929: 613-614.

Oratemnus indicus (With): Beier, 1932b: 591; Beier, 1932e: 60; Roewer, 1936: figs 44a-b; Roewer, 1937: 284; Beier, 1967g: 353; Beier, 1973d: 49; Murthy and Ananthakrishnan, 1977: 126; Sivaraman, 1981: 22; Chandrashekhar, Suryanarayanan and Murthy, 1990: 108; Harvey, 1991a: 467; Beron, 2002: 36.

Not Atemnus indicus (With): Beier, 1930e: 297-298 (misidentification; see Anatemnus nilgiricus Beier).

 

Type locality: Vellore, near Chennai (as Madras), Tamil Nadu, India.

Distribution: India; Sri Lanka.

Oratemnus loyolai Sivaraman

Oratemnus loyolai Sivaraman, 1980c: 349-352, figs 3a-b; Harvey, 1991a: 467; Klausen, 2005: 645, fig. 15.

 

Type locality: Red Hills, Chennai (as Madras), Tamil Nadu, India.

Distribution: India; Sri Lanka.

Oratemnus manilanus Beier

Oratemnus manilanus Beier, 1932b: 590, fig. 14; Beier, 1932e: 59, fig. 70; Roewer, 1937: 284; Harvey, 1991a: 467.

 

Type locality: Manila, Manila, Luzon Island, Philippines.

Distribution: Philippines.

Oratemnus navigator (With)

Chelifer politus Simon: Hansen, 1884: 543 (misidentification).

Chelifer navigator With, 1906: 191-193, fig. 22, plate 4 fig. 9a; With, 1907: 62-63, figs 15-16; Ellingsen, 1909a: 219; Ellingsen, 1911b: 141; Ellingsen, 1914: 3-4.

Oratemnus navigator (With): Beier, 1932b: 589-590; Beier, 1932e: 58-59; Roewer, 1937: 284; Beier, 1948b: 462; Murthy and Ananthakrishnan, 1977: 126-127; Harvey, 1991a: 467; Klausen, 2005: 645-646, fig. 16; Paul, Donovan and Sudarsanam, 2005: 70.

Not Chelifer (Atemnus) navigator With: Ellingsen, 1910a: 358 (misidentification; see Oratemnus timorensis Beier).

 

Type locality: on ship from India.

Distribution: India; Indonesia (Bali, Java); Malaysia; Myanmar; Philippines; Seychelles; Sri Lanka; Thailand.

Oratemnus philippinensis Beier

Oratemnus philippinensis Beier, 1932b: 592-593; Beier, 1932e: 61, fig. 74; Roewer, 1937: 284; Beier, 1966b: 342-343; Harvey, 1991a: 467.

 

Type locality: Los Baños, Luzon Island, Philippines.

Distribution: Philippines.

Oratemnus proximus Beier

Oratemnus proximus Beier, 1932b: 590-591; Beier, 1932e: 59-60, fig. 71; Roewer, 1937: 284; Beier, 1973d: 49-50; Harvey, 1991a: 467.

 

Type locality: Sumatra, Indonesia.

Distribution: Indonesia (Sumatra); Sri Lanka.

Oratemnus punctatus (L. Koch and Keyserling)

Chelifer punctatus L. Koch and Keyserling, 1885: 45-46, plate 4 figs 3, 3a-c.

Chelifer brevidigitatus L. Koch and Keyserling, 1885: 48-49, plate 4 figs 6, 6a-c; With, 1905: 112, 328; Rainbow, 1916: 34; Judson, 1997a: 11-12 (as junior synonym of Oratemnus punctatus (L. Koch and Keyserling)) (synonymised by Beier, 1948c: 529).

Chelifer (Atemnus) brevidigitatus L. Koch and Keyserling: Ellingsen, 1910a: 357.

Anatemnus brevidigitatus (L. Koch and Keyserling): Beier, 1932b: 586-587; Beier, 1932e: 56, fig. 67; Chamberlin, 1934b: 8; Roewer, 1937: 284.

Anatemnus (?) punctatus (L. Koch and Keyserling): Beier, 1932e: 56; Chamberlin, 1934b: 8; Roewer, 1937: 284.

Steiratemnus punctatus (L. Koch and Keyserling): Beier, 1948c: 529-531, figs 2-3.

Oratemnus punctatus (L. Koch and Keyserling): Beier, 1966a: 296; Beier, 1969a: 187; Beier, 1975b: 203; Beier, 1976f: 213; Harvey, 1981b: 245; Harvey, 1985b: 128-129; Harvey, 1991a: 467-468; Judson, 1997a: 36; Klausen, 2005: 646, fig. 17.

Not Chelifer brevidigitatus L. Koch and Keyserling: Ellingsen, 1912a: 121 (misidentification; true identity not known).

 

Type locality: of Chelifer punctatus: Gayndah, Queensland, Australia.

of Chelifer brevidigitatus: Rockhampton, Queensland, Australia.

Distribution: Australia (Australian Capital Territory, Lord Howe Island; New South Wales, Queensland, South Australia, Victoria, Western Australia).

Oratemnus saigonensis (Beier)

Chelifer (Atemnus) saigonensis Beier, 1930d: 197-198, figs 1a-b.

Anatemnus saigonensis (Beier): Beier, 1932b: 583; Beier, 1932e: 53, fig. 65; Roewer, 1937: 283.

Oratemnus timorensis Beier: Redikorzev, 1938: 99 (misidentification).

Oratemnus saigonensis (Beier): Beier, 1951a: 84-86, fig. 23; Beier, 1967g: 353-354; Chapman, 1984a: 6; Harvey, 1991a: 468; Jędryczkowski, 1998: 187.

 

Type locality: Ho Chi Minh (as Saigon), Ho Chi Minh, Vietnam.

Distribution: Cambodia; Laos; Malaysia (Sarawak); Thailand; Vietnam.

Oratemnus samoanus Beier

Oratemnus samoanus Beier, 1932b: 593, fig. 16; Beier, 1932e: 61-62, fig. 75; Chamberlin, 1934b: 8; Roewer, 1937: 284; Chamberlin, 1939b: 212-215, figs 3a-m; Beier, 1966d: 140; Beier, 1970g: 320; Beier, 1982: 43; Harvey, 1991a: 468.

Oratemnus samoanus samoanus Beier: Beier, 1957d: 30-31.

 

Type locality: Apia, Upolu, Samoa.

Distribution: Federated States of Micronesia; French Polynesia (Îles Marquises); Guam; Jamaica; Japan; Marshall Islands; Northern Mariana Islands; Papua New Guinea; Samoa; Solomon Islands; St Kitts and Nevis.

Oratemnus samoanus samoanus Beier

Oratemnus samoanus samoanus Beier: Beier, 1957d: 30-31.

 

Distribution: French Polynesia (Îles Marquises); Jamaica; Marshall Islands; Papua New Guinea; Samoa; Solomon Islands; St Kitts and Nevis.

Oratemnus samoanus whartoni Chamberlin

Oratemnus whartoni Chamberlin, 1947b: 308-312, figs 2a-e.

Oratemnus samoanus whartoni Chamberlin: Beier, 1957d: 30, fig. 15; Morikawa, 1960: 138-139; Samuelson and Nishida, 1987: 169; Harvey, 1991a: 468; Sato, 1994b: 80.

 

Type locality: Ylig Bay, Guam.

Distribution: Federated States of Micronesia; Guam; Japan; Marshall Islands; Northern Mariana Islands.

Oratemnus semidivisus Redikorzev

Oratemnus semidivisus Redikorzev, 1938: 99-100, figs 27-28; Roewer, 1940: 346; Beier, 1951a: 86; Harvey, 1991a: 468; Schawaller, 1994: 745; Jędryczkowski, 1998: 187.

 

Type locality: Da Lat (as Dalat), Lâm Dong, Vietnam; Mt Hon-Ba, Vietnam; and Phsar Ream (as Réam), Kampot, Cambodia.

Distribution: Cambodia; Thailand; Vietnam.

Oratemnus timorensis Beier

Chelifer (Atemnus) navigator With: Ellingsen, 1910a: 358 (misidentification).

Oratemnus timorensis Beier, 1932b: 591; Beier, 1932e: 60, fig. 72; Roewer, 1937: 284; Petersen, 1968: 120; Harvey, 1991a: 468.

Not Oratemnus timorensis Beier: Redikorzev, 1938: 99 (misidentification; see Oratemnus saigonensis (Beier)).

 

Type locality: Telang, Timor, Indonesia.

Distribution: Indonesia (Timor); Solomon Islands.

Oratemnus yodai Morikawa

Oratemnus yodai Morikawa, 1968: 262-263, figs 1c, 2a, 2f; Harvey, 1991a: 469.

 

Type locality: Kapure, near Mt Numbur, Nepal.

Distribution: Nepal.

Genus Paratemnoides Harvey

Paratemnus Beier, 1932b: 562-563; Beier, 1932e: 33; Morikawa, 1960: 137; Murthy and Ananthakrishnan, 1977: 125; Sivaraman, 1980c: 345-346 (junior homonym of Paratemnus Ameghino, 1904).

Paratemnoides Harvey, 1991a: 469 (replacement name for Paratemnus Beier, 1932b).

 

Type species: Chelifer guineensis Ellingsen, 1906 (junior synonym of Chelifer pallidus Balzan, 1892), by original designation.

Distribution: AFRICA: widespread.

ASIA: widespread.

AMERICAS: widespread.

Paratemnoides aequatorialis (Beier)

Chelifer (Atemnus) guineensis Ellingsen, 1906: 246 (misidentification, in part; see Paratemnoides guineensis (Ellingsen)).

Paratemnus aequatorialis Beier, 1932b: 565-566; Beier, 1932e: 35, fig. 35; Roewer, 1937: 282.

Paratemnoides aequatorialis (Beier): Harvey, 1991a: 469.

 

Type locality: Guinea-Bissau.

Distribution: Guinea-Bissau.

Paratemnoides assimilis (Beier)

Chelifer birmanicus Thorell: Tullgren, 1912a: 267 (misidentification); Tullgren, 1912b: 269 (misidentification); Dammerman, 1922: 100 (misidentification); Dammerman, 1929: 114 (not seen) (misidentification); Bristowe, 1931b: 1390 (misidentification); Dammerman, 1948: 496 (misidentification).

Paratemnus assimilis Beier, 1932b: 569-570, fig. 9; Beier, 1932e: 40, fig. 45; Roewer, 1937: 283; Harvey, 1988b: 338-341, figs 86-95.

Paratemnoides assimilis (Beier): Harvey, 1991a: 469.

 

Type locality: Kolambugan, Lanao del Norte, Mindanao, Philippines.

Distribution: Indonesia (Java, Krakatau Islands); Philippines.

Paratemnoides borneoensis (Beier)

Paratemnus borneoensis Beier, 1932b: 571; Beier, 1932e: 41, fig. 46; Roewer, 1937: 283; Weidner, 1959: 116.

Paratemnoides borneoensis (Beier): Harvey, 1991a: 469.

 

Type locality: Lebang Hara, Kalimantan, Indonesia.

Distribution: Indonesia (Kalimantan).

Paratemnoides curtulus (Redikorzev)

Anatemnus curtulus Redikorzev, 1938: 91-93, figs 19-21; Roewer, 1940: 346.

Paratemnus curtulus (Redikorzev): Beier, 1951a: 78; Beier, 1967g: 352.

Paratemnoides curtulus (Redikorzev): Harvey, 1991a: 469; Jędryczkowski, 1998: 187.

 

Type locality: Tram Hanh (as Arbré-Broyé), Lâm Dong, Vietnam; Da Lat (as Dalat), Lâm Dong, Vietnam; Côn So’n (as Poulo-Condore), Khanh Hoa, Vietnam; Pleiku, Gai Lai, Vietnam; Angkor, Cambodia.

Distribution: Cambodia; Vietnam.

Paratemnoides ellingseni (Beier)

Chelifer (Atemnus) feae Ellingsen: Ellingsen, 1910a: 357-358 (misidentification, in part).

Chelifer feae Ellingsen: Ellingsen, 1912b: 79, 90-91 (misidentification, in part); Ellingsen, 1913a: 452 (misidentification); Godfrey, 1927: 18 (misidentification).

Paratemnus ellingseni Beier, 1932b: 572; Beier, 1932e: 42-43, fig. 50; Lawrence, 1937: 268; Roewer, 1937: 283; Caporiacco, 1941: 45; Beier, 1958a: 173; Beier, 1964k: 71-72; Beier, 1966k: 464; Beier, 1967d: 80; Lawrence, 1967: 89; Mahnert, 1983b: 369-371, figs 30-34; Mahnert, 1984b: 64.

Paratemnoides ellingseni (Beier): Harvey, 1991a: 470; Klausen, 2005: 646, figs 4, 22.

 

Type locality: Baía de Maputo (as Delagoa Bay), Maputo, Mozambique.

Distribution: Democratic Republic of Congo (Zaire); Ethiopia; Ghana; Kenya; Madagascar; Mozambique; Somalia; South Africa; Uganda; Zimbabwe.

Paratemnoides elongatus (Banks)

Atemnus elongatus Banks, 1895a: 10; Banks, 1904a: 141; Coolidge, 1908: 112; Banks, 1909c: 174; Chamberlin, 1931a: fig. 11k.

Atemnus floridanus Tullgren, 1900a: 153-155, plate 1 figs 1-4 (synonymised by Banks, 1904a: 141).

Lustrochernes (?) floridanus (Tullgren): Beier, 1932e: 95; Roewer, 1937: 290; Hoff, 1958: 21.

Genus ? elongatus Banks: Beier, 1932e: 277; Roewer, 1937: 317.

Paratemnus elongatus (Banks): Hoff, 1946b: 109-113, figs 1-2; Hoff, 1958: 20; Hoff, 1964b: 2-5; Lloyd and Muchmore, 1974: 381; Muchmore, 1974b: 80; Lloyd, Correale and Muchmore, 1975: 241-242, fig. 1; Muchmore, 1975d: fig. 13; Beier, 1976d: 46; Muchmore, 1977: 76; Brach, 1978: 4-11, figs 1-4; Brach, 1979: 34; Corey and Stout, 1990: 169-171; Muchmore, 1990a: 517, fig. 18.59; J.A. Zeh and D.W. Zeh, 1990: 308-310, fig. 1; Muchmore, 1991b: 157; dos Santos and Coutinho-Netto, 2006: 28-32.

Paratemnoides elongatus (Banks): Harvey, 1991a: 470; Poinar, Ćurčić and Cokendolpher, 1998: 86.

Not Atemnus elongatus Banks: Berger, 1906: 489-490, fig. 1 (misidentification; see Paratemnoides nidificator (Balzan)).

 

Type locality: of Atemnus elongatus: St Lucie River, Sand Point, Enterprise, Punta Gorda, Volusia County, Florida, U.S.A.

of Atemnus floridanus: Victoria, Lake County, Florida, U.S.A.; and Apopka, Orange County, Florida, U.S.A.

Distribution: Cuba; Dominican Republic; Guatemala; Mexico; Panama; U.S.A. (Alabama, Florida); U.S. Virgin Islands.

Paratemnoides feai (Ellingsen)

Chelifer (Atemnus) feae Ellingsen, 1906: 246-248; Ellingsen, 1910a: 357-358 (in part; see Paratemnoides ellingseni).

Chelifer feae Ellingsen: Ellingsen, 1912b: 79, 90-91 (in part; see Paratemnoides ellingseni).

Paratemnus feae (Ellingsen): Beier, 1932b: 571-572; Beier, 1932e: 42, figs 33, 49; Roewer, 1937: 283.

Paratemnus feai (Ellingsen): Vachon, 1956: 2-3.

Paratemnoides feai (Ellingsen): Harvey, 1991a: 470.

Not Chelifer feae Ellingsen: Ellingsen, 1913a: 452 (misidentification; see Paratemnoides ellingseni); Godfrey, 1927: 18 (misidentification; see Paratemnoides ellingseni).

 

Type locality: San Thiago, near Praia, Praia, Cape Verde.

Distribution: Cameroon; Cape Verde.

Paratemnoides guianensis (Caporiacco)

Paratemnus guianensis Caporiacco, 1947b: 21; Caporiacco, 1948a: 617-618, fig. 11.

Paratemnoides guianensis (Caporiacco): Harvey, 1991a: 470.

 

Type locality: Linden (as MacKenzie), Demerara-Mahaica, Guyana.

Distribution: Guyana.

Paratemnoides indicus (Sivaraman)

Paratemnus indicus Sivaraman, 1980c: 346-347, figs 1a-b; Sivaraman, 1982: 187-194, figs 1a-e, 4.

Paratemnoides indicus (Sivaraman): Harvey, 1991a: 471.

 

Type locality: Pondicherry, Pondicherry, India.

Distribution: India.

Paratemnoides indivisus (Tullgren)

Chelifer indivisus Tullgren, 1907d: 7-8, figs 5a-c; Ellingsen, 1912b: 79.

Cyclatemnus indivisus (Tullgren): Beier, 1932b: 562.

Cyclatemnus (?) indivisus (Tullgren): Beier, 1932e: 32; Roewer, 1937: 282.

Paratemnus indivisus (Tullgren): Mahnert, 1983b: 372-375, figs 40-47.

Paratemnoides indivisus (Tullgren): Harvey, 1991a: 471.

 

Type locality: Mombo, Tanga, Tanzania.

Distribution: Kenya; Tanzania.

Paratemnoides insubidus (Tullgren)

Chelifer insubidus Tullgren, 1907a: 58-59, figs 17a-b; Tullgren, 1908b: 285, fig. 3; Ellingsen, 1912b: 79, 91.

Paratemnus insubidus (Tullgren): Beier, 1932b: 566; Beier, 1932e: 36, fig. 36; Roewer, 1937: 282; Beier, 1947b: 324; Weidner, 1959: 114; Beier, 1964k: 71; Beier, 1966k: 464.

Paratemnoides insubidus (Tullgren): Harvey, 1991a: 471; Klausen, 2005: 646.

 

Type locality: Port Elizabeth, Eastern Cape, South Africa.

Distribution: Namibia; South Africa.

Paratemnoides insularis (Banks)

Atemnus insularis Banks, 1902a: 68-69, plate 2 fig. 11; Snodgrass, 1902: 80; Roth and Craig, 1970: 119; Baert, Maelfait and Desender, 1995: 16.

Paratemnus insularis (Banks): Chamberlin, 1934b: 8.

Paratemnoides insularis (Banks): Harvey, 1991a: 471.

 

Type locality: Isla Isabela (as Albemarle Island), Galapagos Islands, Galápagos, Ecuador.

Distribution: Ecuador (Galapagos Islands).

Paratemnoides japonicus (Morikawa)

Paratemnus japonicus Morikawa, 1953a: 348-350, figs 1h-i, 2a-b (as Paratenmus [sic] japonicus); Morikawa, 1960: 138, plate 3 fig. 6, plate 8 figs 1, 16, plate 9 fig. 17; Morikawa, 1962: 419, fig. 4; Sato, 1994b: 80.

Paratemnoides japonicus (Morikawa): Harvey, 1991a: 471.

 

Type locality: Ishite River, Matsuyama, Ehime Prefecture, Shikoku, Japan.

Distribution: Japan.

Paratemnoides laosanus (Beier)

Paratemnus laosanus Beier, 1951a: 76-78, fig. 19; Beier, 1967g: 352.

Paratemnoides laosanus (Beier): Harvey, 1991a: 471.

 

Type locality: Vieng Chang, Mekong, Laos.

Distribution: India; Laos.

Paratemnoides magnificus (Beier)

Atemnus rotundus (With): Beier, 1930a: 207-208, fig. 5 (misidentification).

Paratemnus magnificus Beier, 1932b: 568-569; Beier, 1932e: 39, fig. 42; Roewer, 1937: 283; Beier, 1965g: 767.

Paratemnoides magnificus (Beier): Harvey, 1991a: 471.

 

Type locality: Hughipagu, Papua New Guinea.

Distribution: Papua New Guinea.

Paratemnoides mahnerti (Sivaraman)

Paratemnus robustus Sivaraman, 1980c: 347-349, figs 2a-b (junior primary homonym of Paratemnus robustus Beier, 1932b).

Paratemnus mahnerti Sivaraman, 1981b: 96 (replacement name for Paratemnus robustus Sivaraman).

Paratemnoides mahnerti (Sivaraman): Harvey, 1991a: 471-472.

 

Type locality: Pondicherry, Pondicherry, India.

Distribution: India.

Paratemnoides minutissimus (Beier)

Paratemnus minutissimus Beier, 1974d: 901-902, fig. 2.

Paratemnoides minutissimus (Beier): Harvey, 1991a: 472.

 

Type locality: Caiobá, Paraná, Brazil.

Distribution: Brazil (Paraná).

Paratemnoides nidificator (Balzan)

Chelifer nidificator Balzan, 1888a: no pagination, figs; Balzan, 1890: 417-418, figs 5, 5a-b; Tullgren, 1907a: 56; With, 1908a: 321-324, text-fig. 83, figs 36a-c.

Chelifer (Atemnus) nidificator minor Balzan, 1892: 510-511, fig. 1; Ellingsen, 1905e: 323-324 (synonymised by Klausen, 2005: 646).

Chelifer (Atemnus) nidificator Balzan: Balzan, 1892: 547; Ellingsen, 1902: 146-148; Ellingsen, 1905c: 2; Ellingsen, 1910a: 358.

Atemnus elongatus Banks: Berger, 1906: 489-490, fig. 1 (misidentification).

Chelifer nidificator minor Balzan: Ellingsen, 1909b: 216.

Lamprochernes nidificator (Balzan): Beier, 1930a: 211.

Paratemnus nidificator (Balzan): Beier, 1932b: 567-568; Beier, 1932e: 37-38, fig. 39; Roewer, 1937: 283; Mello-Leitão, 1939a: 118; Mello-Leitão, 1939b: 615; Feio, 1945: 5; Hoff, 1946c: 3; Schenkel, 1953a: 57; Muchmore, 1971a: 82.

Paratemnus minor (Balzan): Beier, 1932b: 568; Beier, 1932e: 38, fig. 40; Roewer, 1937: 283; Feio, 1945: 5; Mello-Leitão and Feio, 1949: 320, plate 4 fig. 1; Beier, 1959e: 203; Mahnert, 1979d: 763; Adis, 1981: 119, etc.; N.S. Hahn and Matthiesen, 1982: 99; Adis and Mahnert, 1986: 311; Mahnert and Adis, 1986: 213; Mahnert, Adis and Bührnheim, 1987: fig. 14a; Hahn and Matthiesen, 1993a: 346-352, figs 1-12; Hahn and Matthiesen, 1993b: 572-574; Muchmore, 1993a: 92.

Paratemnoides minor (Balzan): Harvey, 1991a: 472; Aguiar and Bührnheim, 1994: 74; Morais, Adis, Mahnert and Berti-Filho, 1997: 481; Aguiar and Bührnheim, 1998a: 384; Aguiar and Bührnheim, 1998b: 453; Mahnert and Adis, 2002: 378; Aguiar and Bührnheim, 2003: 517; Ceballos and Florez, 2007: 49.

Paratemnoides nidificator (Balzan): Harvey, 1991a: 472; Poinar, Ćurčić and Cokendolpher, 1998: 90; Tizo-Pedroso and Del-Claro, 2005: 873-877, figs 1-4; Klausen, 2005: 646-647, fig. 19; Tizo-Pedroso and Del-Claro, 2007: 124-130, figs 1-4.

 

Type locality: of Chelifer nidificator: Asunción, Central, Paraguay; Rio Apa, Paraguay; Mato Grosso (as Matto-grosso), Brazil.

of Chelifer (Atemnus) nidificator minor: San Estéban, Venezuela.

Distribution: Argentina; Brazil (Amazonas, Bahia, Distrito Federal, Mato Grosso, Pará, Paraná, Roraima, São Paulo); Colombia; Costa Rica; Ecuador; Guatemala; Guyana; Haiti; Mexico; Panama; Paraguay; Peru; St Vincent and the Grenadines; Venezuela.

Paratemnoides obscurus (Beier)

Paratemnus obscurus Beier, 1959d: 42-43, fig. 20.

Paratemnoides obscurus (Beier): Harvey, 1991a: 472.

 

Type locality: Lubumbashi (as Elisabethville), Shaba, Democratic Republic of Congo (Zaire).

Distribution: Democratic Republic of Congo (Zaire).

Paratemnoides pallidus (Balzan)

Chelifer (Atemnus) pallidus Balzan, 1892: 511-512, figs 2, 2a (incorrectly claimed to be a junior homonym of Chernes pallidus Banks, 1890, by Ellingsen, 1912b: 79).

Chelifer (Atemnus) guineensis Ellingsen, 1906: 246 (in part; see Paratemnoides aequatorialis (Beier)); Ellingsen, 1910a: 358 (synonymised by Harvey, 1991a: 472).

Chelifer plebejus With: Tullgren, 1907a: 55, fig. 15 (misidentification); Ellingsen, 1914: 4 (misidentification, in part).

Chelifer guineensis Ellingsen: Ellingsen, 1912b: 79.

Atemnus plebejus (With): Beier, 1930a: 208 (misidentification).

Paratemnus guineensis (Ellingsen): Beier, 1932b: 565; Beier, 1932e: 34-35, fig. 34; Roewer, 1937: 282; Vachon, 1952a: 30; Beier, 1972a: 11; Majer, 1974: 90; Mahnert, 1978f: 97-101, figs 56-68; Heurtault, 1983: 18; Mahnert, 1983b: 369.

Paratemnus congicus Beier, 1932b: 566-567, fig. 7; Beier, 1932e: 36-37, figs 37-38; Roewer, 1936: fig. 83a; Roewer, 1937: 282; Beier, 1948b: 452, 457; Beier, 1954d: 134; Beier, 1959d: 42; Beier, 1967d: 80 (synonymised by Beier, 1972a: 11).

Paratemnus ceylonicus Beier, 1932b: 569, fig. 8; Beier, 1932e: 39-40, figs 43-44; Roewer, 1937: 283; Beier, 1973d: 47; Murthy and Ananthakrishnan, 1977: 126 (synonymised by Klausen, 2005: 647).

Paratemnoides ceylonicus (Beier): Harvey, 1991a: 469.

Paratemnoides pallidus (Balzan): Harvey, 1991a: 472-473; Poinar, Ćurčić and Cokendolpher, 1998: 85; Klausen, 2005: 647, fig. 18.

 

Type locality: of Chelifer (Atemnus) pallidus: Sierra Leone.

of Chelifer (Atemnus) guineensis: Rio Cassine, Guinea-Bissau; Ribeira Palma, São Tomé (as Thomé), São Tomé and Príncipe; Basilé, Fernando Poo (now Macias Nguema), Equatorial Guinea; Punta Frailes, Fernando Poo (now Macias Nguema), Equatorial Guinea; and Lake N'Komi (as Fernand-Vaz), Ogooué-Maritime, Gabon.

of Paratemnus congicus: Eala, Équateur, Democratic Republic of Congo (Zaire).

of Paratemnus ceylonicus: Sri Lanka (as Ceylon).

Distribution: Burundi; Cameroon; Congo; Côte d’Ivoire; Democratic Republic of Congo (Zaire); Equatorial Guinea; Gabon; Guinea; Guinea-Bissau; India; Kenya; Malaysia; São Tomé and Príncipe; Sierra Leone; Sri Lanka; Togo; Uganda.

Paratemnoides perpusillus (Beier)

Paratemnus perpusillus Beier, 1935a: 487-488, fig. 3; Roewer, 1937: 283; Beier, 1948b: 457; Muchmore, 1971a: 87.

Paratemnoides perpusillus (Beier): Harvey, 1991a: 473; Judson, 1997a: 34.

 

Type locality: Barbados.

Distribution: Barbados.

Paratemnoides persimilis (Beier)

Paratemnus persimilis Beier, 1932b: 567; Beier, 1932e: 37; Roewer, 1937: 283; Vachon, 1952a: 30, fig. 18; Beier, 1965b: 373.

Paratemnoides persimilis (Beier): Harvey, 1991a: 473.

 

Type locality: Edéa, Littoral, Cameroon.

Distribution: Cameroon; Chad; Guinea.

Paratemnoides philippinus (Beier)

Paratemnus philippinus Beier, 1932b: 573, fig. 10; Beier, 1932e: 43-44, fig. 52; Roewer, 1937: 283; Beier, 1957d: 28; Morikawa, 1960: 138; Beier, 1966b: 342; Morikawa, 1960: 138; Sato, 1991b: 218; Sato, 1994b: 80.

Paratemnoides philippinus (Beier): Harvey, 1991a: 473.

 

Type locality: Vinac Island, Philippines.

Distribution: Japan; Philippines.

Paratemnoides plebejus (With)

Chelifer plebejus With, 1906: 185-188, figs 18-20, plate 4 figs 7a-b; Ellingsen, 1910a: 358; Ellingsen, 1911a: 36; Ellingsen, 1911b: 142; Ellingsen, 1914: 4 (in part; see Paratemnoides ceylonicus (Beier)).

Paratemnus plebejus (With): Beier, 1932b: 570; Beier, 1932e: 40; Roewer, 1937: 283; Beier, 1952a: 105-106, figs 7-8; Beier, 1955e: 43.

Paratemnoides plebejus (With): Harvey, 1991a: 473; Judson, 1997a: 34-35.

Not Chelifer plebejus With: Tullgren, 1907a: 55, fig. 15 (misidentification; see Paratemnoides ceylonicus (Beier)).

Not Atemnus plebejus (With): Beier, 1930a: 208 (misidentification; see Paratemnoides ceylonicus (Beier)).

 

Type locality: Car Nicobar (as Kar-Nicobar), Andaman and Nicobar Islands, India.

Distribution: Myanmar; India (Nicobar Islands); Indonesia (Sumatra); Malaysia; Myanmar; Papua New Guinea; Singapore.

Paratemnoides pococki (With)

Chelifer javanus Thorell: Pocock, 1900: 156 (misidentification).

Chelifer pococki With, 1907: 63-65, figs 17-19.

Paratemnus pococki (With): Beier, 1932b: 574; Beier, 1932e: 45; Roewer, 1937: 283; Harvey, 1985b: 129.

Paratemnoides pococki (With): Harvey, 1991a: 473-474; Judson, 1997a: 35.

 

Type locality: north coast of Christmas Island, Australia.

Distribution: Australia (Christmas Island).

Paratemnoides redikorzevi (Beier)

Anatemnus robustus Redikorzev, 1938: 93-95, figs 22, 25a; Roewer, 1940: 346 (junior secondary homonym of Paratemnus robustus Beier, 1932b).

Paratemnus redikorzevi Beier, 1951a: 74-76, fig. 18; Beier, 1967g: 352 (replacement name for Anatemnus robustus Redikorzev).

Paratemnoides redikorzevi (Beier): Harvey, 1991a: 474; Schawaller, 1994: 745-747, figs 44-45; Jędryczkowski, 1998: 188.

 

Type locality: Kontum, Vietnam.

Distribution: Thailand; Vietnam.

Paratemnoides robustus (Beier)

Paratemnus robustus Beier, 1932b: 572-573; Beier, 1932e: 43, fig. 51; Roewer, 1937: 283; Beier, 1966b: 342.

Paratemnoides robustus (Beier): Harvey, 1991a: 474.

 

Type locality: Paniol, Manila, Philippines.

Distribution: Philippines.

Paratemnoides salomonis (Beier)

Paratemnus salomonis Beier, 1935e: 639-640, fig. 2; Roewer, 1937: 283; Beier, 1957d: 28-29, figs 14a-c; Beier, 1964f: 594; Beier, 1965g: 767; Beier, 1966d: 140; Beier, 1970g: 320; Beier, 1982: 43; Hammen, 1983b: 17, 29-30.

Paratemnus salomonis papuanus Beier, 1948c: 556-558, fig. 19; Judson, 1997a: 33 (as junior synonym of Paratemnoides salomonis (Beier)) (synonymised by Beier, 1965g: 767).

Paratemnoides salomonis (Beier): Harvey, 1991a: 474; Judson, 1997a: 38; Klausen, 2005: 647-648, fig. 20.

 

Type locality: of Paratemnus salomonis: Rendova Island, Western, Solomon Islands.

of Paratemnus salomonis papuanus: Kokoda, Northern, Papua New Guinea.

Distribution: Federated States of Micronesia; Indonesia (West Papua); Northern Mariana Islands; Papua New Guinea; Solomon Islands; Vanuatu.

Paratemnoides salomonis hebridicus (Beier)

Paratemnus salomonis hebridicus Beier, 1940a: 174-175, fig. 3; Beier, 1966d: 140.

Paratemnoides salomonis hebridicus (Beier): Harvey, 1991a: 474; Judson, 1997a: 19.

 

Type locality: Ounua, Malekula, Vanuatu.

Distribution: Solomon Islands; Vanuatu.

Paratemnoides salomonis salomonis (Beier)

 

Distribution: Federated States of Micronesia; Indonesia (West Papua); Northern Mariana Islands; Papua New Guinea; Solomon Islands.

Paratemnoides sinensis (Beier)

Paratemnus sinensis Beier, 1932b: 573-574, fig. 11; Beier, 1932e: 44, fig. 53; Roewer, 1937: 283.

Paratemnoides sinensis (Beier): Harvey, 1991a: 474; Schawaller, 1995b: 1057.

 

Type locality: near Guangzhou (as Canton), Guangdong, China.

Distribution: China (Guangdong).

Paratemnoides singularis (Beier)

Paratemnus singularis Beier, 1965g: 767-768, fig. 9; Tenorio and Muchmore, 1982: 378.

Paratemnoides singularis (Beier): Harvey, 1991a: 474.

 

Type locality: Port Moresby, National Capital, Papua New Guinea.

Distribution: Papua New Guinea.

Paratemnoides sumatranus (Beier)

Paratemnus sumatranus Beier, 1935a: 485-487, fig. 2; Roewer, 1937: 283.

Paratemnoides sumatranus (Beier): Harvey, 1991a: 474.

 

Type locality: Singkarak (as Singkarah), Sumatera Berat, Sumatra, Indonesia.

Distribution: Indonesia (Sumatra).

Genus Progonatemnus Beier

Progonatemnus Beier, 1955g: 51-52; Harvey, 1991a: 474.

 

Type species: Progonatemnus succineus Beier, 1955g, by original designation.

Progonatemnus succineus Beier

Progonatemnus succineus Beier, 1955g: 52-53, fig. 3; Schawaller, 1978: 4; Harvey, 1991a: 475; Selden, 1993b: 305.

 

Type locality: Baltic Amber (Eocene).

Distribution: Baltic Amber (Eocene).

 

CATALOGUE

 

Family Atemnidae Chamberlin 1931

         Subfamily Atemninae Chamberlin 1931

                  Genera Anatemnus to Progonatemnus