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Pseudoscorpions of the World |
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Last updated: 19 January 2009 |
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Version 1.2, 2009 |


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Family CHTHONIIDAE Daday Chthoniidae Daday: Hansen, 1894: 232; Tullgren, 1906a: 205; With, 1906: 64; Kew, 1911a: 55; Lessert, 1911: 37; Chamberlin, 1929a: 56; Chamberlin, 1931a: 211-212; Beier, 1932a: 23; Beier, 1932g: 181-182; Chamberlin, 1935b: 479 (as Chthonidae [sic]); Roewer, 1937: 235-236; Hoff, 1949b: 431; Hoff, 1951: 1; G.O. Evans and Browning, 1954: 7; Petrunkevitch, 1955: 80; Hoff, 1956a: 2; Hoff, 1959a: 7; Morikawa, 1960: 93; Dubinin, 1962: 441; Meinertz, 1962: 40-41; Beier, 1963b: 17-18; Murthy and Ananthakrishnan, 1977: 11; Muchmore, 1982a: 97; Harvey, 1985b: 137; Legg, 1987: 179; Legg and Jones, 1988: 55; Harvey, 1991a: 131; Gärdenfors and Wilander, 1992: 28; Harvey, 1992c: 1399. Cecodithinae Chamberlin and R.V. Chamberlin, 1945: 65 (as Cecodithiinae [sic]) (synonymised by Judson, 2001b: 141). Subfamily CHTHONIINAE Daday Chthoniinae Daday, 1888: 133; Balzan, 1892: 545; Hansen, 1894: 232; With, 1906: 64-66; Lessert, 1911: 37; Redikorzev, 1924b: 26; Chamberlin, 1929a: 62-63; Chamberlin, 1931a: 212; Beier, 1932a: 35; Morikawa, 1960: 93. Tribe CHTHONIINI Daday Kewochthoniini Chamberlin, 1929a: 63 (as Kewochthonini [sic]). Chthoniini Daday: Chamberlin, 1929a: 68 (as Chthonini [sic]); Chamberlin, 1931a: 212; Beier, 1932a: 35; Roewer, 1937: 236; Hoff, 1951: 2; Hoff, 1956a: 2; Hoff, 1959a: 7. Genus Anisoditha Chamberlin and R.V. Chamberlin Anisoditha Chamberlin and R.V. Chamberlin, 1945: 26; Harvey, 1991a: 218; Dashdamirov and Judson, 2004: 49-50.
Type species: Chthonius curvidigitatus Balzan, 1887b, by original designation. Distribution: SOUTH AMERICA: Paraguay. Anisoditha curvidigitata (Balzan) Chthonius curvidigitatus Balzan, 1887b: no pagination, figs; Balzan, 1890: 449-450, figs 27, 27a; Balzan, 1892: 550. Chthonius (Sigmodactylus) curvidigitatus Balzan: Hadži, 1930b: 140. Verrucaditha (?) curvidigitata (Balzan): Beier, 1932a: 32; Roewer, 1937: 235. Anisoditha curvidigitata (Balzan): Chamberlin and R.V. Chamberlin, 1945: 26-27; Harvey, 1991a: 218.
Type locality: Rio Apa, Paraguay. Distribution: Paraguay. Genus Aphrastochthonius Chamberlin Aphrastochthonius Chamberlin, 1962: 307-308; Muchmore, 1972f: 433-435; Muchmore, 1973b: 47; Muchmore, 1986b: 17; Harvey, 1991a: 134; Muchmore, 1992: 127-129.
Type species: Aphrastochthonius tenax Chamberlin, 1962, by original designation. Distribution: NORTH AMERICA: Cuba; Guatemala; Mexico; U.S.A. Aphrastochthonius alteriae Muchmore Aphrastochthonius alteriae Muchmore, 1977: 64, figs 1-3; Muchmore, 1982f: 63, fig. 1; Harvey, 1991a: 134; Muchmore, 1991b: 155; Ceballos, 2004: 427; Villegas-Guzman, 2006: 133.
Type locality: Group IV, Ruinas de Palenque, Chiapas, Mexico. Distribution: Mexico. Aphrastochthonius cubanus Dumitresco and Orghidan Aphrastochthonius cubanus Dumitresco and Orghidan, 1977: 99-101, figs 1a-g; Harvey, 1991a: 135.
Type locality: Isla de Pinos, Isla de la Juventud, Cuba. Distribution: Cuba. Aphrastochthonius grubbsi Muchmore Aphrastochthonius grubbsi Muchmore, 1984f: 171-173, figs 1-2; Harvey, 1991a: 135.
Type locality: Lost Piton Cave, 6.5 km SE. of Angels Camp, Calaveras County, California, U.S.A. Distribution: U.S.A. (California). Aphrastochthonius major Muchmore Aphrastochthonius major Muchmore, 1973b: 47-48, fig. 1; Reddell and Elliott, 1973b: 183; Reddell, 1981: 118-119, fig. 18; Harvey, 1991a: 135; Ceballos, 2004: 427.
Type locality: Cueva de la Capilla, 13.5 km NW. of Gómez Farías, Tamaulipas, Mexico. Distribution: Mexico. Aphrastochthonius pachysetus Muchmore Aphrastochthonius pachysetus Muchmore, 1976a: 361-363, figs 1-3; Harvey, 1991a: 135; Muchmore, 1992: 129.
Type locality: Doc Brito Cave, 4.5 miles NE. of Whites City, Eddy County, New Mexico, U.S.A. Distribution: U.S.A. (New Mexico). Aphrastochthonius palmitensis Muchmore Aphrastochthonius palmitensis Muchmore, 1986b: 17-19, fig. 1; Harvey, 1991a: 135; Ceballos, 2004: 427.
Type locality: Grutas del Palmito, 7 km SSW. of Bustamante, Nuevo León, Mexico. Distribution: Mexico. Aphrastochthonius parvus Muchmore Aphrastochthonius parvus Muchmore, 1972f: 437-438, figs 4-7; Reddell and Elliott, 1973a: 173; Reddell, 1981: 119, fig. 18; Harvey, 1991a: 135; Ceballos, 2004: 427.
Type locality: La Cueva de la Florida, 15 km SSW. of Mante, Tamaulipas, Mexico. Distribution: Mexico. Aphrastochthonius patei Muchmore Aphrastochthonius patei Muchmore, 1982f: 63-64, fig. 2; Harvey, 1991a: 135; Ceballos, 2004: 427.
Type locality: Sumidero de Oyamel (as Cueva de Oyamel), Sistema Purificación, SW. of El Barretal, Tamaulipas, Mexico. Distribution: Mexico. Aphrastochthonius pecki Muchmore Aphrastochthonius pecki Muchmore, 1968a: 17-18, figs 1-2; Peck, 1989: 20; Harvey, 1991a: 135.
Type locality: Crystal Caverns, Clay, Jefferson County, Alabama, U.S.A. Distribution: U.S.A. (Alabama). Aphrastochthonius russelli Muchmore Aphrastochthonius russelli Muchmore, 1972f: 440-441, figs 10-12; Reddell and Elliott, 1973a: 173; Reddell, 1981: 119, fig. 18; Harvey, 1991a: 135; Ceballos, 2004: 427.
Type locality: La Cueva Pinta, about 12 km NE. of Valles, San Luis Potosí, Mexico. Distribution: Mexico. Aphrastochthonius similis Muchmore Aphrastochthonius similis Muchmore, 1984f: 173-174, fig. 3; Drewry, 1989: 566; Harvey, 1991a: 136.
Type locality: Carlow's Cave, 7.5 km SE. of Angels Camp, Calaveras County, California, U.S.A. Distribution: U.S.A. (California). Aphrastochthonius tenax Chamberlin Aphrastochthonius tenax Chamberlin, 1962: 308-310, figs 2a-j; Muchmore, 1972f: 435, figs 1-3; Peck, 1989: 20; Harvey, 1991a: 136.
Type locality: Bangor Cave, near Bangor, Blount County, Alabama, U.S.A. Distribution: U.S.A. (Alabama). Aphrastochthonius verapazanus Muchmore Aphrastochthonius verapazanus Muchmore, 1972f: 438-440, figs 8-9; Muchmore, 1977: 64; Reddell, 1981: 119, fig. 18; Harvey, 1991a: 136.
Type locality: La Cueva Sepacuite #2, Senahu Finca Sepacuite, Alta Verapaz, Guatemala. Distribution: Guatemala. Genus Apochthonius Chamberlin Apochthonius Chamberlin, 1929a: 66-67; Chamberlin, 1929e: 152; Beier, 1932a: 41; Hoff, 1949b: 434; Hoff, 1956a: 2; Benedict and Malcolm, 1973: 621; Muchmore and Benedict, 1976: 68-69; Harvey, 1991a: 136; Muchmore, 1992: 129. Apochthonius (Apochthonius) Chamberlin: Beier, 1932a: 41.
Type species: Chthonius moestus Banks, 1891, by original designation. Distribution: NORTH AMERICA: Canada; U.S.A. Apochthonius coecus (Packard) Chthonius coecus Packard, 1884: 203; Packard, 1888: 46-48, plate 11 figs 4, 4a-c; Banks, 1895a: 13; Coolidge, 1908: 114; Roewer, 1937: 239; Wolf, 1938: 606. Chthonius (?) coecus Packard: Beier, 1932a: 61; Hoff, 1958: 4. Apochthonius coecus (Packard): Muchmore, 1963a: 9-11, figs 5-6; Holsinger, 1963: 34; Holsinger, 1964: 63; Holsinger and Culver, 1988: 42, fig. 15; Harvey, 1991a: 136.
Type locality: Weyer's Cave, Augusta County, Virginia, U.S.A. Distribution: U.S.A. (Virginia). Apochthonius colecampi Muchmore Apochthonius colecampi Muchmore, 1967d: 89-90, fig. 1; Craig, 1975: 3; Muchmore, 1976e: 76; Harvey, 1991a: 136; Elliott, 2007: 142.
Type locality: Cole Camp Cave, SE. of Lincoln, Benton County, Missouri, U.S.A. Distribution: U.S.A. (Missouri). Apochthonius diabolus Muchmore Apochthonius diabolus Muchmore, 1967d: 92, fig. 3; Harvey, 1991a: 136.
Type locality: Devil's Den, Devil's Den State Park, Washington County, Arkansas, U.S.A. Distribution: U.S.A. (Arkansas). Apochthonius forbesi Benedict Apochthonius forbesi Benedict, 1979: 79-82, figs 1-3; Harvey, 1991a: 136.
Type locality: 14 km S., 11 km E. of Bend, Deschutes County, Oregon, U.S.A. Distribution: U.S.A. (Oregon). Apochthonius grubbsi Muchmore Apochthonius grubbsi Muchmore, 1980b: 93-94, figs 1-3; Drewry, 1989: 566; Harvey, 1991a: 137.
Type locality: Music Hall Cave, 4.5 miles SE. of Angels Camp, Calaveras County, California, U.S.A. Distribution: U.S.A. (California). Apochthonius hobbsi Muchmore Apochthonius sp.: Hobbs and Flynn, 1981: 12 (not seen). Apochthonius hobbsi Muchmore, 1994b: 317-319, fig. 1.
Type locality: Buckskin Cave I, Ross County, Ohio, U.S.A. Distribution: U.S.A. (Ohio). Apochthonius holsingeri Muchmore Apochthonius holsingeri Muchmore, 1967d: 93-94, fig. 5; Muchmore, 1976e: 77; Holsinger and Culver, 1988: 42, fig. 15; Harvey, 1991a: 137.
Type locality: Cave Run Pit Cave, about 8 miles N. of Warm Springs, Bath County, Virginia, U.S.A. Distribution: U.S.A. (Virginia). Apochthonius hypogeus Muchmore Apochthonius hypogeus Muchmore, 1976e: 75-76, fig. 8; Harvey, 1991a: 137.
Type locality: Great North Mountain, Augusta County, Virginia, U.S.A. Distribution: U.S.A. (Virginia). Apochthonius indianensis Muchmore Apochthonius indianensis Muchmore, 1967d: 92-93, fig. 4; Muchmore, 1976e: 76; Harvey, 1991a: 137; Lewis, 1994: 15; McCreedy, 2004: 180, 205.
Type locality: Donaldson's Cave, Spring Mill State Park, Lawrence County, Indiana, U.S.A. Distribution: U.S.A. (Indiana). Apochthonius intermedius Chamberlin Apochthonius (Apochthonius) intermedius Chamberlin, 1929e: 153; Beier, 1932a: 42. Apochthonius intermedius Chamberlin: Chamberlin, 1931a: figs 22a-b, 46a; Beier, 1932g: fig. 223; Roewer, 1937: 238; Snodgrass, 1948: fig. 12a; Vachon, 1949: fig. 239; Hoff, 1958: 6; R.O. Schuster, 1966b: figs 4, 8-9; Harvey, 1991a: 137.
Type locality: Friday Harbour, San Juan Island, San Juan County, Washington, U.S.A. Distribution: U.S.A. (Washington). Apochthonius irwini R.O. Schuster Apochthonius irwini R.O. Schuster, 1966b: 179-182, figs 11-12; Harvey, 1991a: 137.
Type locality: 5 miles NW. of Spanish Flat, Napa County, California, U.S.A. Distribution: U.S.A. (California). Apochthonius knowltoni Muchmore Apochthonius knowltoni Muchmore, 1980a: 87-89, figs 1-5; Harvey, 1991a: 137.
Type locality: 10 miles SE. of Smoot, Lincoln County, Wyoming, U.S.A. Distribution: U.S.A. (Wyoming). Apochthonius magnanimus Hoff Apochthonius magnanimus Hoff, 1956a: 4-10, figs 2-4; Hoff, 1958: 6; Hoff, 1959b: 4, etc.; Welbourn, 1978: 31; D.W. Zeh, 1987b: 1086; Harvey, 1991a: 137; Muchmore, 1992: 129.
Type locality: W. of Cowles, San Miguel County, New Mexico, U.S.A. Distribution: U.S.A. (New Mexico). Apochthonius malheuri Benedict and Malcolm Apochthonius malheuri Benedict and Malcolm, 1973: 621-626, figs 1-6; Drewry, 1989: 566; Harvey, 1991a: 138. Apochthonius sp.: Peck, 1973: 100.
Type locality: Malheur Cave, 13 miles SE. of Princeton, Harney County, Oregon, U.S.A. Distribution: U.S.A. (Oregon). Apochthonius maximus R.O. Schuster Apochthonius maximus R.O. Schuster, 1966b: 182, figs 3, 10, 13; Harvey, 1991a: 138.
Type locality: Caspar Loop Road, Russian Gulch State Park, Mendocino County, California, U.S.A. Distribution: U.S.A. (California). Apochthonius minimus R.O. Schuster Apochthonius minimus R.O. Schuster, 1966b: 178-179, figs 1, 2, 5, 7; D.L. Johnson and Wellington, 1980a: 339-349; D.L. Johnson and Wellington, 1980b: 353-362; Harvey, 1991a: 138; Witt and Dill, 1996: 396-405, fig. 2; Buddle, 2005: 15.
Type locality: 2 miles N. of Brinnon, Olympic Peninsula, Jefferson County, Washington, U.S.A. Distribution: Canada (British Columbia); U.S.A. (Oregon, Washington). Apochthonius minor Muchmore Apochthonius sp.: Holsinger and Peck, 1971: 30. Apochthonius minor Muchmore, 1976e: 72-75, figs 6-7; Harvey, 1991a: 138.
Type locality: Parker Cave, 2 miles NE. of Subligna, Chatooga County, Georgia, U.S.A. Distribution: U.S.A. (Georgia). Apochthonius moestus (Banks) Chthonius moestus Banks, 1891: 165; Banks, 1895a: 13; Coolidge, 1908: 114; Ewing, 1911: 80. Apochthonius moestus (Banks): Chamberlin, 1929a: 67, figs 2c, 3g (in part); Chamberlin, 1931a: figs 6a, 21f; Beier, 1932a: 41-42, figs 47-48 (in part); Roewer, 1937: 238; Hoff, 1944b: 125, figs 1c-d; Hoff, 1945a: 105-109, figs 4-7; Pearse, 1946: 257; Hoff, 1958: 6 (in part); Muchmore and Benedict, 1976: 69-73, figs 1-9; Rowland and Reddell, 1976: 14; Stryker and Taylor, R.W., 1977: 26; Sweeney, Taylor and Counts, 1977: 55; Taylor, Sweeney and Counts, 1977: 115; D.W. Zeh, 1987b: 1086; Harvey, 1991a: 138; Snider and Nelson, 1991: 260; Buddle, 2005: 15. Apochthonius sp. prob. moestus (Banks): Graves and Graves, 1969: 267. Not Apochthonius moestus (Banks): Chamberlin, 1929a: 67 (in part) (misidentification; true identification unknown); Beier, 1930a: 202 (misidentification; true identification unknown); Hoff, 1945i: 311-313 (misidentification; true identification unknown); Hoff, 1949b, figs 11, 15a-d: 434 (misidentification; true identification unknown); Hoff, 1951: 4-5 (misidentification; true identification unknown); Hoff, 1952a: 42-43 (misidentification; true identification unknown); Hoff, 1956a: 2-4, fig. 1 (misidentification; true identification unknown); Hoff and Bolsterli, 1956: 158-159 (misidentification; true identification unknown); Hoff, 1958: 6 (in part) (misidentification; true identification unknown); Hoff, 1959b: 4, etc. (misidentification; true identification unknown); Nelson, 1975: 279-280, figs 14-17 (misidentification; true identification unknown). Not Apochthonius (Apochthonius) moestus (Banks): Chamberlin, 1929e: 153 (misidentification; true identification unknown); Beier, 1932a: 41-42 (in part) (misidentification; true identification unknown).
Type locality: Ithaca, Tompkins County, New York, U.S.A. Distribution: Canada (Ontario); U.S.A. (Indiana, Illinois, Michigan, Missouri, New Mexico, New York, North Carolina, Pennsylvania, West Virginia). Apochthonius mysterius Muchmore Apochthonius sp.: Lewis, 1974: 9. Apochthonius mysterius Muchmore, 1976e: 70, fig. 4; Peck and Lewis, 1977: 47; Harvey, 1991a: 138; Elliott, 2007: 142, 150.
Type locality: Red Fork Passage, Mystery Cave, about 5 miles SE. of Perryville, Perry County, Missouri, U.S.A. Distribution: U.S.A. (Missouri). Apochthonius occidentalis Chamberlin Apochthonius occidentalis Chamberlin, 1929a: 67-68; Beier, 1932a: 42; Roewer, 1937: 238; Hoff, 1958: 6; Harvey, 1991a: 139.
Type locality: Portland, Multnomah County, Oregon, U.S.A. Distribution: U.S.A. (Oregon). Apochthonius paucispinosus Muchmore Apochthonius paucispinosus Muchmore, 1967d: 90-92, fig. 2; Harvey, 1991a: 139; Drewry, 1989: 566.
Type locality: Bennett Cave, about 2 miles N. of Dryfork, Tucker County, West Virginia, U.S.A. Distribution: U.S.A. (West Virginia). Apochthonius russelli Muchmore Apochthonius russelli Muchmore, 1976e: 70-72, fig. 5; Peck, 1989: 20, fig. 6; Harvey, 1991a: 139.
Type locality: Pig Entrance, Russell Cave, Russell Cave National Monument, Jackson County, Alabama, U.S.A. Distribution: U.S.A. (Alabama). Apochthonius titanicus Muchmore Apochthonius titanicus Muchmore, 1976e: 67-69, figs 1-3; Harvey, 1991a: 139.
Type locality: near 'The Titans', Blanchard Springs Caverns, 3 miles E. of Fifty Six, Stone County, Arkansas, U.S.A. Distribution: U.S.A. (Arkansas). Apochthonius typhlus Muchmore Apochthonius typhlus Muchmore, 1967d: 94-95, fig. 6; Craig, 1975: 3; Harvey, 1991a: 139; Elliott, 2007: 142.
Type locality: Old Spanish Cave, Stone County, Missouri, U.S.A. Distribution: U.S.A. (Missouri). Genus Austrochthonius Chamberlin Austrochthonius Chamberlin, 1929a: 68; Beier, 1932a: 38; Beier, 1976f: 203; Harvey, 1991a: 139; Harvey, 1991e: 487-488; Harvey, 1996a: 256; Judson, 2001b: 141-142. Paraustrochthonius Beier, 1931b: 52; Beier, 1932a: 40 (synonymised by Beier, 1976f: 203). Cecoditha Mello-Leitão, 1939a: 115-116; Chamberlin and R.V. Chamberlin, 1945: 65-66; Harvey, 1991a: 218 (synonymised by Judson, 2001b: 142).
Type species: of Austrochthonius: Chthonius chilensis Chamberlin, 1923b, by original designation. of Paraustrochthonius: Paraustrochthonius tullgreni Beier, 1931b, by original designation. of Cecoditha: Cecoditha parva Mello-Leitão, 1939a, by original designation.
Distribution: AFRICA: South Africa. AUSTRALASIA: Australia; New Zealand. SOUTH AMERICA: Argentina; Bolivia; Brazil; Chile; Paraguay. INDIAN OCEAN: Îles Crozet. Austrochthonius argentinae Hoff Austrochthonius argentinae Hoff, 1950b: 225-228, figs 1-6; Beier, 1974d: 899; Harvey, 1991a: 139.
Type locality: Pampa de Olaen, Córdoba, Argentina. Distribution: Argentina; Brazil (Santa Catarina). Austrochthonius australis Hoff Austrochthonius australis Hoff, 1951: 5-9, figs 1-3; Beier, 1966a: 281-283, fig. 2; Beier, 1969a: 187; Dartnall, 1970: 65-66; Beier, 1975b: 203; Hickman and Hill, 1978: appendix a; Koch and Majer, 1980: 25; Harvey, 1981b: 239; Main, 1981: 1065; Harvey, 1985b: 137; C.M.A. Kennedy, 1990b: 234-239, figs 1a-f, 2g-l, 3m-q, 4s-w; Eberhard, Richardson and Swain, 1991: 26; Harvey, 1991a: 139; Harvey, 1992c: figs 8-13; Harvey, 1994b: 156; Austin, Gibson and Harvey, 1998: 346, 349.
Type locality: Mt Slide, Victoria, Australia. Distribution: Australia (Australian Capital Territory, New South Wales, Queensland, Tasmania, Victoria, Western Australia). Austrochthonius bolivianus Beier Austrochthonius bolivianus Beier, 1930a: 201-202; Beier, 1932a: 38-39, fig. 42; Roewer, 1937: 238; Beier, 1974d: 899; Harvey, 1991a: 140.
Type locality: Bolivia. Distribution: Bolivia; Brazil (Santa Catarina). Austrochthonius cavicola Beier Austrochthonius cavicola Beier, 1968a: 757-758, fig. 1; Harvey, 1981b: 239; D.C. Lee, 1983: 183; Harvey, 1991a: 140; Moulds, 2004: 12.
Type locality: Cathedral Cave, Naracoorte, South Australia, Australia. Distribution: Australia (South Australia). Austrochthonius chilensis (Chamberlin) Chthonius chilensis Chamberlin, 1923b: 187-189, figs 20a-j. Austrochthonius chilensis (Chamberlin): Chamberlin, 1929a: 68; Chamberlin, 1931a: fig. 21e; Beier, 1932a: 38; Roewer, 1937: 238; Beier, 1962h: 131; Beier, 1964b: 312-313; Cekalovic, 1984: 8; Harvey, 1991a: 140.
Type locality: Butalcura, Isla Grande Chiloé, Los Lagos, Chile. Distribution: Argentina; Chile. Austrochthonius chilensis chilensis (Chamberlin) Distribution: Chile. Austrochthonius chilensis magalhanicus Beier Austrochthonius chilensis magalhanicus Beier, 1964b: 313; Cekalovic, 1976: 17; Cekalovic, 1984: 9; Harvey, 1991a: 140.
Type locality: Magallanes y Antártica Chilena (as Magellanes), Chile. Distribution: Chile. Austrochthonius chilensis transversus Beier Austrochthonius chilensis transversus Beier, 1964j: 488, fig. 1; Harvey, 1991a: 140.
Type locality: Mt Piltriquitron, El Bolsón, Río Negro, Argentina. Distribution: Argentina. Austrochthonius easti Harvey Austrochthonius easti Harvey, 1991e: 488-490, figs 1-7; Harvey, Gray, Hunt and Lee, 1993: 130, 131.
Type locality: Dry Swallett Cave, C-18, Cape Range peninsula, Western Australia, Australia. Distribution: Australia (Western Australia). Austrochthonius iguazuensis Vitali-di Castri Austrochthonius iguazuensis Vitali-di Castri, 1975: 117-119, figs 1-2, 5-6, 9-10; Harvey, 1991a: 140.
Type locality: Cataratas del Iguazú, Paraná, Brazil. Distribution: Brazil (Paraná). Austrochthonius insularis Vitali-di Castri Austrochthonius insularis Vitali-di Castri, 1968: 141-144, figs 1-7; Harvey, 1991a: 140.
Type locality: Île de la Possession, Îles Crozet, French Southern and Antarctic Lands. Distribution: French Southern and Antarctic Lands (Îles Crozet). Austrochthonius mordax Beier Austrochthonius inversus mordax Beier, 1967b: 282-283, fig. 4; Palma, Lovis and Tither, 1989: 20. Austrochthonius mordax Beier: Beier, 1976f: 203-204; Harvey, 1991a: 140.
Type locality: Lake Wairarapa, Wellington, North Island, New Zealand. Distribution: New Zealand. Austrochthonius muchmorei Harvey and Mould Chthonius caecus Tullgren, 1909b: 414-415, fig. 3 (junior primary homonym of Chthonius coecus Packard, 1884, and Chthonius caecus Simon, 1885b). Mundochthonius (?) caecus (Tullgren): Beier, 1932a: 38; Roewer, 1937: 238. Mundochthonius caecus (Tullgren): Nicholls, 1933: 111; Chamberlin, 1934b: 3; Weidner, 1959: 115. Sathrochthonius tullgreni Chamberlin, 1962: 306-307 (replacement name for Chthonius caecus Tullgren); Harvey, 1981b: 241; Harvey, 1985b: 140-141; Harvey, 1991a: 202 (junior secondary homonym of Austrochthonius tullgreni (Beier)). Sathrochthonius (?) tullgreni Chamberlin: Beier, 1966a: 276; Muchmore, 1982d: 158. Austrochthonius muchmorei Harvey and Mould, 2006: 209-210 (replacement name for Sathrochthonius tullgreni Chamberlin).
Type locality: Brunswick Junction (as Brunswick), Western Australia, Australia. Distribution: Australia (Western Australia). Austrochthonius paraguayensis Vitali-di Castri Austrochthonius paraguayensis Vitali-di Castri, 1975: 121-124, figs 3-4, 7-8; Harvey, 1991a: 140.
Type locality: Ciudad del Este (as Puerto Presidente Stroessner), Alto Parana, Paraguay. Distribution: Paraguay. Austrochthonius parvus (Mello-Leitão) Cecoditha parva Mello-Leitão, 1939a: 116-117, figs 1a-b; Chamberlin and R.V. Chamberlin, 1945: 66-67, figs 17a-b; Harvey, 1991a: 218; Kury and Noguiera, 1999: 13. Austrochthonius parvus (Mello-Leitão): Judson, 2001b: 142-144, figs 1-9.
Type locality: Puerto Madryn (as Madryn), Chubut, Argentina. Distribution: Argentina. Austrochthonius persimilis Beier Austrochthonius persimilis Beier, 1930d: 205-206, figs 10a-d (as Austrochtonius [sic] persimilis); Beier, 1932a: 39, fig. 43; Roewer, 1937: 238; Beier, 1964b: 313-314, fig. 1; Cekalovic, 1984: 9; Harvey, 1991a: 141.
Type locality: Pitrufquén, Araucanía, Chile. Distribution: Chile. Austrochthonius rapax Beier Austrochthonius rapax Beier, 1976f: 204, fig. 4; Harvey, 1991a: 141.
Type locality: Tuna Saddle, Taumarunui, North Island, New Zealand. Distribution: New Zealand. Austrochthonius semiserratus Beier Austrochthonius semiserratus Beier, 1930d: 206-207, figs 11a-b, 12-13; Beier, 1932a: 39, figs 44-45; Roewer, 1937: 238; Beier, 1974d: 899; Vitali-di Castri and Castri, 1976: 62, 63 (as Austrochthonius semisseratus [sic]); Cekalovic, 1984: 9; Harvey, 1991a: 141. Not Austrochthonius semiserratus Beier: Beier, 1964b: 314 (misidentification; see Chiliochthonius montanus Vitali-di Castri).
Type locality: Temuco, Araucanía, Chile. Distribution: Brazil (Santa Catarina); Chile. Austrochthonius strigosus Harvey and Mould Austrochthonius strigosus Harvey and Mould, 2006: 206-209, figs 1-5.
Type locality: Ludlow region (site LDMB2, 33.58921503°S, 115.4904651°E), Western Australia, Australia. Distribution: Australia (Western Australia). Austrochthonius tullgreni (Beier) Paraustrochthonius tullgreni Beier, 1931b: 53; Beier, 1932a: 40, fig. 46; Roewer, 1937: 238; Beier, 1958a: 159; Beier, 1964k: 31; Beier, 1966k: 455-456; Lawrence, 1967: 87. Austrochthonius tullgreni (Beier): Harvey, 1991a: 141.
Type locality: Pietermaritzburg, KwaZulu-Natal, South Africa. Distribution: South Africa. Austrochthonius zealandicus Beier Austrochthonius zealandicus Beier, 1966c: 363-364, fig. 1; Anonymous, 1970: 24; Tenorio and Muchmore, 1982: 384; Harvey, 1991a: 141. Austrochthonius luxtoni Beier, 1967b: 279-280, fig. 2; Palma, Lovis and Tither, 1989: 20 (synonymised by Beier, 1976f: 203). Austrochthonius inversus Beier, 1967b: 280-282, fig. 3; Beier, 1969e: 413, 414; Wise, 1970: 234; Palma, Lovis and Tither, 1989: 19 (synonymised by Beier, 1976f: 203).
Type locality: of Austrochthonius zealandicus: Birdlings Flat, Canterbury, South Island, New Zealand. of Austrochthonius luxtoni: Mackenzie's Bush, near Oparau, South Auckland, New Zealand. of Austrochthonius inversus: Palmerston North, Pohangina Valley, Silverstream, Wellington, North Island, New Zealand. Distribution: New Zealand. Austrochthonius zealandicus obscurus Beier Austrochthonius obscurus Beier, 1966c: 364-365, fig. 2; Beier, 1967b: 279; Tenorio and Muchmore, 1982: 382. Austrochthonius zealandicus obscurus Beier: Beier, 1976f: 203; Harvey, 1991a: 141.
Type locality: Scenic Reserve, Kaituna Valley, Banks Peninsula, Canterbury, South Island, New Zealand. Distribution: New Zealand. Austrochthonius zealandicus zealandicus Beier Austrochthonius zealandicus zealandicus Beier: Beier, 1976f: 203.
Distribution: New Zealand. Genus Caribchthonius Muchmore Caribchthonius Muchmore, 1976c: 361-363; Harvey, 1991a: 142.
Type species: Caribchthonius butleri Muchmore, 1976c, by original designation. Distribution: AMERICAS: Belize; U.S. Virgin Islands. Caribchthonius butleri Muchmore Caribchthonius butleri Muchmore, 1976c: 363-365, figs 1-7; Harvey, 1991a: 142; Muchmore, 1993b: 32.
Type locality: Reef Bay Trail, Virgin Islands National Park, St John Island, U.S. Virgin Islands. Distribution: U.S. Virgin Islands. Caribchthonius orthodentatus Muchmore Caribchthonius orthodentatus Muchmore, 1976c: 365-367, figs 8-12; Harvey, 1991a: 142; Muchmore, 1991b: 155 (as Caribchthonius orthodactrylus [sic])
Type locality: Corozal, Corozal, Belize. Distribution: Belize. Genus Chiliochthonius Vitali-di Castri Chiliochthonius Vitali-di Castri, 1976: 1277-1278; Harvey, 1991a: 142.
Type species: Chiliochthonius centralis Vitali-di Castri, 1976, by original designation. Distribution: SOUTH AMERICA: Chile. Chiliochthonius centralis Vitali-di Castri Chiliochthonius centralis Vitali-di Castri, 1976: 1278-1280, figs 1-6; Cekalovic, 1984: 9; Harvey, 1991a: 142.
Type locality: Quebrada La Plata, Santiago, Región Metropolitana, Chile. Distribution: Chile. Chiliochthonius montanus Vitali-di Castri Austrochthonius semiserratus Beier: Beier, 1964b: 314 (misidentification). Chiliochthonius montanus Vitali-di Castri, 1976: 1280-1283, figs 7-12, plate 2; Cekalovic, 1984: 10; Harvey, 1991a: 142.
Type locality: Farellones, Santiago, Región Metropolitana, Chile. Distribution: Chile. |
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CATALOGUE
Family Chthoniidae Daday 1888 Subfamily Chthoniinae Daday 1888 Tribe Chthoniini Daday 1888 Genera Anisoditha to Chiliochthonius |