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Pseudoscorpions of the World |
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Last updated: 19 January 2009 |
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Version 1.2, 2009 |
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Withiidae Chamberlin: Weygoldt, 1970a: 253; Muchmore, 1982a: 101-102; Harvey, 1985b: 154; Harvey, 1991a: 640; Gärdenfors and Wilander, 1992: 29; Harvey, 1992c: 1425. Subfamily PARAGONIOCHERNETINAE Beier Paragoniochernetinae Beier, 1944: 198. Genus Cyrtowithius Beier Cyrtowithius Beier, 1955l: 314-316; Harvey, 1991a: 642.
Type species: Cyrtowithius capensis Beier, 1955l, by original designation. Distribution: AFRICA: southern Africa. Cyrtowithius capensis Beier Chelifer tumuliferus Tullgren: Ellingsen, 1912b: 85, 100-101 (misidentification). Cyrtowithius capensis Beier, 1955l: 316-317, figs 30-31; Harvey, 1991a: 642.
Type locality: Yzerfontein, Western Cape, South Africa. Distribution: South Africa. Cyrtowithius tumuliferus (Tullgren) Chelifer tumuliferus Tullgren, 1908b: 283-284, fig. 1. Caffrowithius tumuliferus (Tullgren): Beier, 1932d: 62; Beier, 1932e: 223; Roewer, 1937: 310. Cyrtowithius tumuliferus (Tullgren): Beier, 1955l: 316; Beier, 1966k: 467; Harvey, 1991a: 642. Not Chelifer tumuliferus Tullgren: Ellingsen, 1912b: 85, 100-101 (misidentification; see Cyrtowithius capensis Beier).
Type locality: Port Nolloth, Northern Cape, South Africa. Distribution: Namibia; South Africa. Genus Ectromachernes Beier Ectromachernes Beier, 1944: 197-198; Harvey, 1991a: 645.
Type species: Ectromachernes mirabilis Beier, 1944, by original designation. Distribution: AFRICA: Africa. Ectromachernes elegans Beier Ectromachernes elegans Beier, 1964k: 77-78, fig. 33; Harvey, 1991a: 645.
Type locality: Coldspring, near Grahamstown, Eastern Cape, South Africa. Distribution: South Africa. Ectromachernes lamottei Vachon Ectromachernes lamottei Vachon, 1952a: 38-40, figs 34-41; Harvey, 1991a: 645.
Type locality: Mont Pierre Richaud (as Mont To), Nimba Mountains, Nzerekore, Guinea. Distribution: Guinea. Ectromachernes mirabilis Beier Ectromachernes mirabilis Beier, 1944: 199-200, fig. 17; Mahnert, 1988a: 41-42, figs 1-4; Harvey, 1991a: 645; Judson, 1997a: 27; Beron, 2002: 38; Harvey, 2004: 442.
Type locality: Jemme Forest (as Djem-Djem-Wald), Ethiopia. Distribution: Ethiopia; Kenya. Ectromachernes rhodesiacus Beier Ectromachernes rhodesiacus Beier, 1964k: 76-77, fig. 32; Harvey, 1991a: 645.
Type locality: Dombashawa, Zimbabwe. Distribution: Zimbabwe. Genus Paragoniochernes Beier Paragoniochernes Beier, 1932e: 189; Beier, 1933a: 548; Harvey, 1991a: 649.
Type species: Chelifer lamellatus Tullgren, 1907b, by original designation. Distribution: AFRICA: southern Africa. Paragoniochernes lamellatus (Tullgren) Chelifer lamellatus Tullgren, 1907b: 222-223, figs 2a-b; Ellingsen, 1912b: 81. Paragoniochernes lamellatus (Tullgren): Beier, 1932e: 189, fig. 197; Roewer, 1937: 304; Beier, 1958a: 178-179, fig. 11; Beier, 1964k: 76; Harvey, 1991a: 650.
Type locality: Van Reenen, Free State, South Africa. Distribution: South Africa. Paragoniochernes parvulus Beier Paragoniochernes parvulus Beier, 1955l: 313-314, fig. 29; Harvey, 1991a: 650.
Type locality: Storms River mouth, Tzitzikama Forest, Eastern Cape, South Africa. Distribution: South Africa. Genus Pseudatemnus Beier Pseudatemnus Beier, 1947b: 330-331; Harvey, 1991a: 654.
Type species: Pseudatemnus lawrencei Beier, 1947b, by original designation. Distribution: AFRICA: southern Africa. Pseudatemnus lawrencei Beier Pseudatemnus lawrencei Beier, 1947b: 331-332, figs 35-36; Harvey, 1991a: 654.
Type locality: River Jonder End, Cape Province, South Africa. Distribution: South Africa. Genus Pseudochernes Beier Pseudochernes Beier, 1954e: 86-87; Harvey, 1991a: 654.
Type species: Pseudochernes crassimanus Beier, 1954e, by original designation. Distribution: AFRICA: east Africa. ASIA: west Asia. Pseudochernes arabicus Mahnert Pseudochernes arabicus Mahnert, 1991: 191-192, figs 55-59.
Type locality: Jīzān, Jīzān, Saudi Arabia. Distribution: Saudi Arabia. Pseudochernes crassimanus Beier Pseudochernes crassimanus Beier, 1954e: 87-88, figs 2-3; Harvey, 1991a: 654.
Type locality: Bwayi (as Wahi), Lake Victoria, Mara, Tanzania. Distribution: Tanzania. Subfamily WITHIINAE Chamberlin Withiinae Chamberlin, 1931c: 290; Beier, 1932e: 192; Roewer, 1937: 305; Morikawa, 1960: 148; Meinertz, 1962: 97; Murthy and Ananthakrishnan, 1977: 145-146; . Tribe CACODEMONIINI Chamberlin Cacodemoniini Chamberlin, 1931c: 292 (as Cacodemonini [sic]); Beier, 1932e: 192-193; Roewer, 1937: 306. Genus Cacodemonius Chamberlin Cacodemonius Chamberlin, 1931c: 292; Beier, 1932e: 193; Harvey, 1991a: 641.
Type species: Withius cactorum Chamberlin, 1923c, by original designation. Distribution: SOUTH AMERICA: widespread. Cacodemonius cactorum (Chamberlin) Withius cactorum Chamberlin, 1923c: 377, plate 1 fig. 7, plate 2 fig. 14; Chamberlin, 1931a: fig. 10f. Cacodemonius cactorum (Chamberlin): Chamberlin, 1931c: 292; Beier, 1932e: 193; Roewer, 1937: 306; Harvey, 1991a: 641; Ceballos, 2004: 429.
Type locality: Isla San Pedro Mártir, Golfo de California, Sonora, Mexico. Distribution: Mexico. Cacodemonius pusillus Beier Cacodemonius pusillus Beier, 1953b: 24, figs 7a-b; Harvey, 1991a: 641.
Type locality: Laguna de Jocotal, San Miguel, El Salvador. Distribution: El Salvador. Cacodemonius quartus Hoff Cacodemonius quartus Hoff, 1946c: 18-21, figs 23-26; Muchmore, 1977: 77; Harvey, 1991a: 641; Muchmore, 1991b: 158; Ceballos, 2004: 429; Villegas-Guzman, 2006: 133.
Type locality: "La Zacualpa", near Villa Comaltitlán, Chiapas, Mexico. Distribution: Mexico. Cacodemonius satanas (With) Chelifer satanas With, 1908a: 245-247, text-fig. 66, figs 8a-c. Cacodemonius satanas (With): Chamberlin, 1931c: 292; Beier, 1932e: 194; Roewer, 1937: 306; Harvey, 1991a: 641; Muchmore, 1993a: 99; Harvey, 2004: 442.
Type locality: Los Trinchéras, Venezuela. Distribution: Venezuela. Cacodemonius segmentidentatus (Balzan) Chelifer segmentidentatus Balzan, 1887b: no pagination, figs; Balzan, 1890: 428-430, figs 13, 13a-b; With, 1908a: 242-245, text-fig. 65, figs 7a-j. Chelifer (Trachychernes) segmentidentatus Balzan: Balzan, 1892: 548; Ellingsen, 1905c: 12-13. Cacodemonius segmentidentatus (Balzan): Chamberlin, 1931c: 292; Beier, 1932e: 193-194; Roewer, 1937: 306; Mello-Leitão, 1939a: 121; Mello-Leitão, 1939b: 617; Beier, 1959e: 215, fig. 24; Beier, 1976d: 55 (as Cacodemonius serratidentatus [sic]); Harvey, 1991a: 642; Muchmore, 1993a: 99; Harvey, 2004: 442.
Type locality: Resistencia, Chaco, Argentina; Rio Apa, Paraguay; and Mato Grosso, Brazil. Distribution: Argentina; Brazil (Mato Grosso); Dominican Republic; Ecuador; Paraguay; Venezuela. Cacodemonius zilchi Beier Cacodemonius zilchi Beier, 1953b: 23-24, figs 10a-c; Harvey, 1991a: 642.
Type locality: Laguna de Zapotitan, La Libertad, El Salvador. Distribution: El Salvador. Tribe JUXTACHELIFERINI Hoff Juxtacheliferini Hoff, 1956d: 22-23. Genus Juxtachelifer Hoff Juxtachelifer Hoff, 1956d: 23; Harvey, 1991a: 509.
Type species: Juxtachelifer fructuosus Hoff, 1956d, by original designation. Distribution: NORTH AMERICA: western U.S.A.; Mexico. Juxtachelifer fructuosus Hoff Juxtachelifer fructuosus Hoff, 1956d: 23-28, figs 11-15; Hoff, 1958: 35; Hoff, 1959b: 5, etc.; Muchmore, 1971a: 91; Muchmore, 1990a: 522; Harvey, 1991a: 509; Harvey, 2004: 442; Villegas-Guzmán and Pérez, 2005a: 65, 67, 76; Francke and Villegas-Guzmán, 2006: 295.
Type locality: Santa Fe, Santa Fe County, New Mexico, U.S.A. Distribution: Mexico; U.S.A. (California, New Mexico). Tribe PROTOWITHIINI Beier Protowithiini Beier, 1955b: 215-216. Genus Protowithius Beier Protowithius Beier, 1955b: 216-217; Harvey, 1991a: 654.
Type species: Protowithius fernandezianus Beier, 1955b, by original designation. Distribution: SOUTH AMERICA: Juan Fernandez Islands Protowithius fernandezianus Beier Protowithius fernandezianus Beier, 1955b: 217-218, fig. 7; Cekalovic, 1984: 28; Harvey, 1991a: 654.
Type locality: Las Chozas, Masatierra, Juan Fernandez Islands, Valparaíso, Chile. Distribution: Chile (Juan Fernandez Islands). Protowithius robustus Beier Protowithius robustus Beier, 1955b: 218-220, fig. 8; Beier, 1957c: 463; Cekalovic, 1984: 28; Harvey, 1991a: 654.
Type locality: Plateau del Yunque, Masatierra, Juan Fernandez Islands, Valparaíso, Chile. Distribution: Chile (Juan Fernandez Islands). Tribe WITHIINI Chamberlin Withiini Chamberlin: Chamberlin, 1931c: 292; Beier, 1932e: 194-195; Roewer, 1937: 306; Morikawa, 1960: 148-149; Murthy and Ananthakrishnan, 1977: 146. Genus Afrowithius Chamberlin Afrowithius Chamberlin, 1931c: 293; Beier, 1932e: 221; Harvey, 1991a: 640.
Type species: Chelifer paradoxus Ellingsen, 1912b, by original designation. Distribution: AFRICA: southern Africa. Afrowithius paradoxus (Ellingsen) Chelifer paradoxus Ellingsen, 1912b: 98-99. Afrowithius paradoxus (Ellingsen): Chamberlin, 1931c: 293; Beier, 1932e: 221; Roewer, 1937: 309; Harvey, 1991a: 640.
Type locality: Ntaba Kandoda, near King William's Town, Eastern Cape, South Africa. Distribution: South Africa. Genus Aisthetowithius Beier Aisthetowithius Beier, 1967d: 83-84; Harvey, 1991a: 640.
Type species: Aisthetowithius rossi Beier, 1967d, by original designation. Distribution: AFRICA: eastern Africa. Aisthetowithius rossi Beier Aisthetowithius rossi Beier, 1967d: 84, fig. 7; Mahnert, 1988a: 49-50, figs 12-14; Harvey, 1991a: 640.
Type locality: Kaimosi Mission, 27 miles NE. of Kisumu, Rift Valley, Kenya. Distribution: Kenya; Tanzania. Genus Balanowithius Beier Balanowithius Beier, 1959e: 223; Harvey, 1991a: 640.
Type species: Balanowithius egregius Beier, 1959e, by original designation. Distribution: SOUTH AMERICA: northern South America. Balanowithius egregius Beier Balanowithius egregius Beier, 1959e: 223-224, fig. 31; Harvey, 1991a: 640; Harvey, 2004: 442.
Type locality: Pichilingue, Los Ríos, Ecuador. Distribution: Ecuador. Balanowithius weyrauchi Beier Balanowithius weyrauchi Beier, 1959e: 224-226, fig. 32; Beier, 1974d: 899; Mahnert, 1975b: fig. 6e; Harvey, 1991a: 641.
Type locality: La Balsa, Rio Canchis, Peru. Distribution: Brazil (Santa Catarina); Peru. † Genus Beierowithius Mahnert Oligowithius Beier, 1937d: 309 (junior homonym of Oligowithius Beier, 1936a). Beierowithius Mahnert, 1979d: 802; Harvey, 1991a: 641 (replacement name for Oligowithius Beier, 1937d).
Type species: Chelifer sieboldtii Menge, 1855, by original designation. Distribution: EUROPE: Baltic Amber. † Beierowithius sieboldtii (Menge) Obisium sieboldtii Menge, in C.L. Koch and Berendt, 1854: 97; Scudder, 1891: 276 Obisium siebodii [sic]). Chelifer sieboldtii (Menge): Menge, 1855: 35-36, plate 5 fig. 10 (Chelifer sieboldii [sic]); Hagen, 1870: 267; Scudder, 1891: 255 (as Chelifer sieboldii [sic]); Beier, 1932e: 278. Oligowithius sieboldtii (Menge): Beier, 1937d: 309, fig. 11; Roewer, 1940: 331; Beier, 1955g: 53 (as Oligowithius sieboldii [sic]); Schawaller, 1979: 4 (as Oligowithius sieboldii [sic]). Beierowithius sieboldtii (Menge): Mahnert, 1979e: 802 (as Beierowithius sieboldii [sic]); Harvey, 1991a: 641.
Type locality: Baltic Amber (Eocene). Distribution: Baltic Amber (Eocene). Genus Cryptowithius Beier Cryptowithius Beier, 1967d: 85-86; Harvey, 1991a: 642.
Type species: Cryptowithius inconspicuus Beier, 1967d, by original designation. Distribution: AFRICA: eastern Africa. Cryptowithius inconspicuus Beier Cryptowithius inconspicuus Beier, 1967d: 86-87, fig. 8; Mahnert, 1988a: 45; Harvey, 1991a: 642.
Type locality: Simu Beach, Kwale, Coast, Kenya. Distribution: Kenya. Genus Cystowithius Harvey Cystowithius Harvey, 2004: 440-441.
Type species: Cystowithius smithersi Harvey, 2004, by original designation. Distribution: SOUTH AMERICA: northern South America north to Mexico. Cystowithius chamberlini Harvey Cystowithius chamberlini Harvey, 2004: 452-454, figs 7, 8, 33-36.
Type locality: Matlalpan, San Luis Potosí, Mexico. Distribution: Guatemala; Mexico. Cystowithius colombicus Harvey Cystowithius colombicus Harvey, 2004: 450-452, figs 7, 8, 28-32; Ceballos and Florez, 2007: 50.
Type locality: Paramo de Monserrate, Cundinamarca, Colombia. Distribution: Colombia. Cystowithius ecuadoricus (Beier) Parawithius (Parawithius) nobilis ecuadoricus Beier, 1959e: 217-218, fig. 26; Harvey, 1991a: 651. Cystowithius ecuadoricus (Beier): Harvey, 2004: 448-450, figs 7, 8, 24-27.
Type locality: Lago Zurucuchu, 11 miles W. of Cuenca, Azuay, Ecuador. Distribution: Ecuador; Peru. Cystowithius smithersi Harvey Cystowithius smithersi Harvey, 2004: 442-448, figs 7-23.
Type locality: Volcán Chiles, ca. 15 km N. of Tulcán, Carchi, Ecuador. Distribution: Ecuador. Genus Dolichowithius Chamberlin Dolichowithius Chamberlin, 1931c: 293; Beier, 1932e: 216; Hoff, 1945h: 1; Harvey, 1991a: 642.
Type species: Chelifer longichelifer Balzan, 1887b, by original designation. Distribution: SOUTH AMERICA: widespread. Subgenus Dolichowithius (Dolichowithius) Chamberlin Dolichowithius (Dolichowithius) Chamberlin: Harvey, 1991a: 643. Dolichowithius (Dolichowithius) argentinus Beier Dolichowithius argentinus Beier, 1959e: 226-227, fig. 33; Harvey, 2004: 442. Dolichowithius (Dolichowithius) argentinus Beier: Harvey, 1991a: 643.
Type locality: Salta, Salta, Argentina. Distribution: Argentina. Dolichowithius (Dolichowithius) brasiliensis (Beier) Withius brasiliensis Beier, 1930a: 218-219, figs 15a-b. Dolichowithius brasiliensis (Beier): Beier, 1932e: 219-220, fig 227; Roewer, 1937: 309. Dolichowithius (Dolichowithius) brasiliensis (Beier): Harvey, 1991a: 643.
Type locality: Rio Grande do Sul, Rio Grande do Sul, Brazil. Distribution: Brazil (Rio Grande do Sul). Dolichowithius (Dolichowithius) canestrinii (Balzan) Chelifer canestrinii Balzan, 1887b: no pagination, figs; Balzan, 1890: 430-431, figs 14, 14a-c; Ellingsen, 1905e: 324; With, 1908a: 236-238, figs 5a-d. Chelifer (Chelifer) canestrinii Balzan: Balzan, 1892: 532, figs 22, 22a; Ellingsen, 1905c: 16-17; Ellingsen, 1910a: 385 (in part; see Parawithius iunctus Beier). Withius canestrinii (Balzan): Chamberlin, 1931a: 168. Dolichowithius canestrinii (Balzan): Chamberlin, 1931c: 293; Beier, 1932e: 220, fig. 228; Roewer, 1937: 309; Mello-Leitão, 1939a: 122; Mello-Leitão, 1939b: 618; Feio, 1945: 8; Caporiacco, 1948a: 618; Beier, 1977c: 112; Harvey, 1991a: 643; Muchmore, 1993a: 98; Muchmore, 1997d: 279; Mahnert and Adis, 2002: 380. Dolichowithius (Dolichowithius) canestrinii (Balzan): Harvey, 1991a: 643.
Type locality: Resistencia, Chaco, Argentina. Distribution: Argentina; Brazil (Bahia, Pará); Ecuador; Guyana; U.S. Virgin Islands; Venezuela. Dolichowithius (Dolichowithius) centralis Beier Dolichowithius centralis Beier, 1953b: 25, fig. 8; Harvey, 1991a: 643. Dolichowithius (Dolichowithius) centralis Beier: Harvey, 1991a: 643.
Type locality: Laguna de Zapotitan, La Libertad, El Salvador. Distribution: El Salvador. Dolichowithius (Dolichowithius) emigrans (Tullgren) Chelifer emigrans Tullgren, 1907a: 35-36, fig. 5. Dolichowithius emigrans (Tullgren): Beier, 1932e: 220-221, fig. 229; Roewer, 1937: 309; Weidner, 1959: 114; Mahnert and Adis, 1986: 214; Mahnert and Adis, 2002: 380. Dolichowithius (Dolichowithius) emigrans (Tullgren): Mahnert, 1979d: 794-795, figs 168-169, 187i; Harvey, 1991a: 643.
Type locality: Manaus (as Manaos), Amazonas, Brazil. Distribution: Brazil (Amazonas). Dolichowithius (Dolichowithius) extensus Beier Chelifer longichelifer Balzan: With, 1908a: 238-242, figs 6a-d (misidentification). Dolichowithius extensus Beier, 1932d: 59-60; Beier, 1932e: 216-217; Roewer, 1937: 309; Beier, 1959e: 226; Muchmore, 1993a: 98; Judson, 1997a: 17; Mahnert and Adis, 2002: 380; Harvey, 2004: 442. Dolichowithius (Dolichowithius) extensus Beier: Harvey, 1991a: 643;
Type locality: La Moka, Venezuela; Riacho del Oro, Gran Chaco, Argentina. Distribution: Argentina; Peru; Venezuela. Dolichowithius (Dolichowithius) granulosus Hoff Dolichowithius granulosus Hoff, 1945h: 3-5, figs 3-5; Harvey, 1991a: 643; Harvey, 2004: 442. Dolichowithius (Dolichowithius) granulosus Hoff: Harvey, 1991a: 643.
Type locality: Essequibo River, Onoro, Uppert Takutu-Upper Essequibo, Guyana. Distribution: Guyana. Dolichowithius (Dolichowithius) intermedius Mahnert Dolichowithius (Dolichowithius) intermedius Mahnert, 1979d: 795-796, figs 160-162, 187m; Harvey, 1991a: 644. Dolichowithius intermedius Mahnert: Adis, 1981: 119, etc.; Adis and Mahnert, 1986: 312; Mahnert and Adis, 1986: 214; Adis, Junk and Penny, 1987: 489; Mahnert and Adis, 2002: 380.
Type locality: Río Tarumã-Mirím, Manaus, Amazonas, Brazil. Distribution: Brazil (Amazonas). Dolichowithius (Dolichowithius) longichelifer (Balzan) Chelifer longichelifer Balzan, 1887b: no pagination, figs; Balzan, 1890: 433-434, figs 16, 16a-c. Chelifer (Chelifer) longichelifer Balzan: Balzan, 1892: 534, fig. 26; Ellingsen, 1905e: 324. Withius longichelifer (Balzan): Chamberlin, 1931a: 168, 170, fig. 30m. Dolichowithius longichelifer (Balzan): Chamberlin, 1931c: 293; Beier, 1932e: 218, fig. 224; Roewer, 1937: 309; Feio, 1945: 7; Muchmore, 1993a: 98. Dolichowithius (Dolichowithius) longichelifer (Balzan): Harvey, 1991a: 644. Dolichowithius (Dolichowithius) aff. longichelifer (Balzan): Aguiar and Bührnheim, 1998b: 456. Not Chelifer longichelifer Balzan: Tullgren, 1907a: 35 (misidentification; see Dolichowithius (Dolichowithius) vicinus Beier); With, 1908a: 238-242, figs 6a-d (misidentification; see Dolichowithius (Dolichowithius) extensus Beier). Not Chelifer (Chelifer) longichelifer Balzan: Ellingsen, 1910a: 386 (misidentification; see Dolichowithius (Dolichowithius) modicus Beier).
Type locality: Resistencia, Chaco, Argentina; Asunción, Central, Paraguay; Rio Apa, Paraguay; Mato Grosso (as Matto-grosso), Brazil. Distribution: Argentina; Brazil (Distrito Federal, Mato Grosso); Ecuador; Paraguay; Venezuela. Dolichowithius (Dolichowithius) mediofasciatus Mahnert Dolichowithius (Dolichowithius) mediofasciatus Mahnert, 1979d: 798-801, figs 170-175; Harvey, 1991a: 644; Aguiar and Bührnheim, 1998b: 456; Mahnert and Adis, 2002: fig. 12c. Dolichowithius mediofasciatus Mahnert: Adis, 1981: 119, etc.; Adis and Mahnert, 1986: 312; Mahnert and Adis, 1986: 214; Adis, Junk and Penny, 1987: 489; Mahnert, Adis, and Bührnheim, 1986: figs 11a, 12c; Aguiar, Vicente da Silva and Bührnheim, 1992: 182-185; Harvey, 1992c: figs 211-214; Aguiar and Bührnheim, 1994: 74; Aguiar, Arruda and Bührnheim, 1997: 83-86, fig. 2; Poinar, Ćurčić and Cokendolpher, 1998: 87; Mahnert and Adis, 2002: 380.
Type locality: Rio Solimões, Manaus, Amazonas, Brazil. Distribution: Brazil (Acre, Amazonas). Dolichowithius (Dolichowithius) minutus Mahnert Dolichowithius (Dolichowithius) minutus Mahnert, 1979d: 796-798, figs 163-167; Harvey, 1991a: 644. Dolichowithius minutus Mahnert: Adis, 1981: 119, etc.; Adis and Mahnert, 1986: 312; Mahnert and Adis, 1986: 214; Adis, Junk and Penny, 1987: 489; Morais, Adis, Mahnert and Berti-Filho, 1997: 481; Mahnert and Adis, 2002: 380.
Type locality: Río Tarumã-Mirím, Manaus, Amazonas, Brazil. Distribution: Brazil (Amazonas). Dolichowithius (Dolichowithius) modicus Beier Chelifer (Chelifer) longichelifer Balzan: Ellingsen, 1910a: 386 (misidentification). Dolichowithius modicus Beier, 1932d: 61; Beier, 1932e: 218- 219, fig. 225; Roewer, 1937: 309. Dolichowithius (Dolichowithius) modicus Beier: Harvey, 1991a: 644.
Type locality: Paraguay. Distribution: Paraguay. Dolichowithius (Dolichowithius) simplex Beier Chelifer (Chelifer) simoni Balzan: Ellingsen, 1910a: 387 (misidentification, in part). Dolichowithius simplex Beier, 1932d: 61; Beier, 1932e: 219, fig. 226; Roewer, 1937: 309; Beier, 1976d: 55; Muchmore, 1997d: 278-279, figs 22-25. Dolichowithius (Dolichowithius) simplex Beier: Harvey, 1991a: 644-645.
Type locality: Puerto Rico. Distribution: Dominican Republic; Puerto Rico. Dolichowithius (Dolichowithius) solitarius Hoff Dolichowithius solitarius Hoff, 1945h: 1-3, figs 1-2. Dolichowithius (Dolichowithius) solitarius Hoff: Harvey, 1991a: 645.
Type locality: Costa Rica. Distribution: Costa Rica. Dolichowithius (Dolichowithius) vicinus Beier Chelifer longichelifer Balzan: Tullgren, 1907a: 35 (misidentification). Dolichowithius vicinus Beier, 1932d: 60-61; Beier, 1932e: 217, fig. 223; Roewer, 1937: 309. Dolichowithius (Dolichowithius) vicinus Beier: Harvey, 1991a: 645.
Type locality: Estancia Postillon, Puerto Max, Paraguay. Distribution: Paraguay. Subgenus Dolichowithius (Oligowithius) Beier Dolichowithius (Oligowithius) Beier, 1936a: 447; Harvey, 1991a: 645.
Type species: Dolichowithius (Oligowithius) abnormis Beier, 1936a, by original designation. Dolichowithius (Oligowithius) abnormis Beier Dolichowithius (Oligowithius) abnormis Beier, 1936a: 446-447, fig. 4; Harvey, 1991a: 645. Dolichowithius abnormis Beier: Roewer, 1940: 347; Muchmore, 1993a: 99.
Type locality: Estado Trujillo, La Ceiba, Venezuela. Distribution: Venezuela. Genus Girardwithius Heurtault Girardwithius Heurtault, 1994a: 198-199.
Type species: Girardwithius pumilus Heurtault, 1994, by original designation. Distribution: AFRICA: western Africa. Girardwithius pumilus Heurtault Girardwithius pumilus Heurtault, 1994a: 199-204, figs 9-15.
Type locality: 10 km S. of Ndouci (as N'Douci), Abidjan, Côte d’Ivoire. Distribution: Côte d’Ivoire; Guinea. Genus Hyperwithius Beier Hyperwithius Beier, 1951a: 99-100; Harvey, 1991a: 646.
Type species: Sundowithius annamensis Redikorzev, 1938, by original designation. Distribution: ASIA: China; Vietnam. Hyperwithius annamensis (Redikorzev) Sundowithius annamensis Redikorzev, 1938: 101-103, figs 29-31; Roewer, 1940: 347. Hyperwithius annamensis (Redikorzev): Beier, 1951a: 100; Harvey, 1991a: 646; Jędryczkowski, 1998: 190.
Type locality: Mt Ba Na (as Bana), Vietnam. Distribution: Vietnam. Hyperwithius dawydoffi Beier Hyperwithius dawydoffi Beier, 1951a: 102-104, figs 34a, 35; Harvey, 1991a: 646; Jędryczkowski, 1998: 190; Harvey, 2004: 442.
Type locality: Núi Lang Bian (as Plateau von Langbian), Lâm Dong, Vietnam. Distribution: Vietnam. Hyperwithius tonkinensis Beier Hyperwithius tonkinensis Beier, 1951a: 100-102, figs 33, 34b; Harvey, 1991a: 646; Schawaller, 1995b: 1061; Jędryczkowski, 1998: 190; Harvey, 2004: 442.
Type locality: Lai Chau (as Lau Chau), Tonkin, Lai Chau, Vietnam. Distribution: China (Sichuan); Vietnam. Genus Metawithius Chamberlin Metawithius Chamberlin, 1931c: 293; Beier, 1932e: 200; Murthy and Ananthakrishnan, 1977: 149; Harvey, 1991a: 646.
Type species: Chelifer murrayi Pocock, 1900, by original designation. Distribution: AUSTRALASIA: widespread Subgenus Metawithius (Metawithius) Chamberlin Metawithius (Metawithius) Chamberlin: Harvey, 1991a: 646. Metawithius (Metawithius) indicus Murthy and Ananthakrishnan Metawithius (Metawithius) indicus Murthy and Ananthakrishnan, 1977: 149-151, figs 47a-b; Harvey, 1991a: 646.
Type locality: Vedanthangal, Tamil Nadu, India. Distribution: India. Metawithius (Metawithius) murrayi (Pocock) Chelifer murrayi Pocock, 1900: 156-157, plate 16 figs 1, 1a; With, 1906: 159-163, plate 3 figs 8a-j; Ellingsen, 1911a: 37; Ellingsen, 1911b: 142. Chelifer (Trachychernes) murrayi Pocock: Ellingsen, 1910a: 376. Withius murrayi (Pocock): Beier, 1930e: 292-293. Metawithius murrayi (Pocock): Chamberlin, 1931c: 293; Beier, 1932e: 200, fig. 205; Roewer, 1936: figs 63b, 65; Roewer, 1937: 308; Harvey, 1985b: 154; Judson, 1997a: 28; Harvey, 2004: 442. Metawithius (Metawithius) murrayi (Pocock): Harvey, 1991a: 646.
Type locality: North West Point, Christmas Island, Australia. Distribution: Australia (Christmas Island); Indonesia (Sumatra, Timor); Myanmar. Metawithius (Metawithius) parvus (Beier) Withius parvus Beier, 1930e: 293-294, fig. 5. Metawithius parvus (Beier): Beier, 1932e: 201, fig. 206; Roewer, 1937: 308. Metawithius (Metawithius) parvus (Beier): Harvey, 1991a: 646.
Type locality: Travancore, Kerala, India. Distribution: India. Metawithius (Metawithius) philippinus Beier Metawithius philippinus Beier, 1937b: 274-275, fig. 5; Roewer, 1940: 347; Harvey, 2004: 442. Metawithius (Metawithius) philippinus Beier: Harvey, 1991a: 647.
Type locality: Limay, Bataan, Luzon Island, Philippines. Distribution: Philippines. Metawithius (Metawithius) spiniventer Redikorzev Metawithius spiniventer Redikorzev, 1938: 103-106, figs 32-34; Roewer, 1940: 347; Beier, 1951a: 98-99, fig. 32; Beier, 1967g: 358; Schawaller, 1994: 757, figs 73-74; Jędryczkowski, 1998: 190; Harvey, 2004: 442. Metawithius (Metawithius) spiniventer Redikorzev: Beier, 1955e: 43; Harvey, 1991a: 647.
Type locality: Mt Hon-Ba, Vietnam; Mt Ba Na (as Bana), Vietnam; and Pouthisat (as Pursat), Pouthisat, Cambodia. Distribution: Cambodia; Indonesia (Sumba); Malaysia; Thailand; Vietnam. Metawithius (Metawithius) spiniventer pauper Beier Metawithius spiniventer pauper Beier, 1953g: 86, fig. 5. Metawithius (Metawithius) spiniventer pauper Beier: Harvey, 1991a: 647.
Type locality: Langgai, Sumba, Nusa Tenggara Timur, Indonesia. Distribution: Indonesia (Sumba). Metawithius (Metawithius) spiniventer spiniventer Redikorzev
Distribution: Cambodia; Malaysia; Thailand; Vietnam. Subgenus Metawithius (Microwithius) Redikorzev Microwithius Redikorzev, 1938: 106. Metawithius (Microwithius) Redikorzev: Beier, 1955e: 45; Harvey, 1991a: 647.
Type species: Microwithius yurii Redikorzev, 1938, by monotypy. Metawithius (Microwithius) bulli Sivaraman Metawithius (Microwithius) bulli Sivaraman, 1980a: 113-115, figs 4a-b; Harvey, 1991a: 647.
Type locality: Bangalore, Karnataka, India. Distribution: India. Metawithius (Microwithius) chamundiensis Sivaraman Metawithius (Microwithius) chamundiensis Sivaraman, 1980a: 111-113, figs 3a-b; Harvey, 1991a: 647.
Type locality: Chamundi Hills, Mysore, Karnataka, India. Distribution: India. Metawithius (Microwithius) tweediei Beier Metawithius (Microwithius) tweediei Beier, 1955e: 43-45, fig. 5; Harvey, 1991a: 647; Harvey, 2004: 442.
Type locality: Telom Valley, near Gunong Siku, Cameron Highlands, Pahang, Malaysia. Distribution: Malaysia (mainland). Metawithius (Microwithius) yurii (Redikorzev) Microwithius yurii Redikorzev, 1938: 106-108, figs 35-38; Roewer, 1940: 347; Beier, 1951a: 104. Metawithius (Microwithius) yurii (Redikorzev): Beier, 1955e: 45; Harvey, 1991a: 647; Harvey, 2004: 442. Metawithius yurii (Redikorzev): Harvey, 1988b: 336-338, figs 79-85; Jędryczkowski, 1998: 191.
Type locality: Duong-Dong, Dao Phú Quoc (as Ile Phu-Quoc), Cambodia; and Côn So’n (as Poulo-Condore), Khanh Hoa, Vietnam. Distribution: Cambodia; Indonesia (Krakatau Islands); Vietnam. Genus Nannowithius Beier Nannowithius Beier, 1932d: 57; Beier, 1932e: 211; Harvey, 1991a: 647-648. Myrmecowithius Beier, 1963f: 195-196 (synonymised by Mahnert, 1988a: 68).
Type species: of Nannowithius: Chelifer aethiopicus Simon, 1900c, by original designation. of Myrmecowithius: Myrmecowithius wahrmani Beier, 1963f, by original designation. Distribution: AFRICA: northern Africa. ASIA: western Asia; Middle East. Nannowithius aethiopicus (Simon) Chelifer aethiopicus Simon, 1900c: 596. Nannowithius aethiopicus (Simon): Beier, 1932d: 57; Beier, 1932e: 211, fig. 217; Roewer, 1937: 309; Mahnert, 1988a: 68, figs 57-59; Harvey, 1991a: 648.
Type locality: Agordat, Gash-Barka, Eritrea. Distribution: Eritrea. Nannowithius buettikeri (Mahnert) Myrmecowithius buettikeri Mahnert, 1980c: 40-42, figs 23-28. Nannowithius buettikeri (Mahnert): Mahnert, 1988a: 68; Harvey, 1991a: 648; Mahnert, 1991: 192.
Type locality: Kushm al Buwaybiyat, Ar Riyād, Saudi Arabia. Distribution: Oman; Saudi Arabia. Nannowithius pakistanicus (Beier) Myrmecowithius pakistanicus Beier, 1978d: 233-234, fig. 2. Nannowithius pakistanicus (Beier): Mahnert, 1988a: 68; Harvey, 1991a: 648.
Type locality: Kohala, Kashmir, Pakistan. Distribution: Pakistan. Nannowithius paradoxus (Mahnert) Myrmecowithius paradoxus Mahnert, 1980c: 38-40, figs 17-22. Nannowithius paradoxus (Mahnert): Mahnert, 1988a: 68; Harvey, 1991a: 648; Mahnert, 1991: 192.
Type locality: Wadi Zabib, Yemen. Zabib (PPL) = 15.52'N, 48.00'E Distribution: Yemen. Nannowithius wahrmani (Beier) Myrmecowithius wahrmani Beier, 1963f: 196-197, fig. 8; Mahnert, 1974c: 384; Mahnert, 1975b: fig. 6d (as Myrmecowithius wahrmanni [sic]). Nannowithius wahrmani (Beier): Mahnert, 1988a: 68; Harvey, 1991a: 648.
Type locality: Wadi Abyad, HaDarom (Southern), Israel. Distribution: Israel. Genus Neowithius Beier Neowithius Beier, 1932d: 62; Beier, 1932e: 223; Harvey, 1991a: 648.
Type species: Chelifer insignis With, 1908a, by original designation. Distribution: SOUTH AMERICA: widespread. Neowithius chilensis (Beier) Chelifer (Chelanops) chilensis Beier, 1930d: 201, figs 4-6. Neowithius chilensis (Beier): Beier, 1932d: 62; Beier, 1932e: 224-225, fig. 233; Roewer, 1937: 309; Harvey, 1991a: 648.
Type locality: Temuco, Araucanía, Chile. Distribution: Chile. Neowithius cubanus (Banks) Chelanops cubanus Banks, 1909c: 173, fig. 3 (as Chelanops cabanus [sic]). Neowithius cubanus (Banks): Hoff, 1947b: 545-547, fig. 37; Harvey, 1991a: 648.
Type locality: Cayamas, Cuba. Distribution: Cuba. Neowithius dubius Beier Chelifer (Trachychernes) brevifemoratus Balzan: Ellingsen, 1910a: 373 (misidentification). Neowithius dubius Beier, 1932d: 62; Beier, 1932e: 224, fig. 232; Roewer, 1937: 310; Feio, 1945: 8; Harvey, 1991a: 649.
Type locality: Theresópolis (now Queçaba), Santa Catarina, Brazil. Distribution: Brazil (Distrito Federal, Santa Catarina). Neowithius exilimanus (Balzan) Chelifer exilimanus Balzan, 1887b: no pagination, figs; Balzan, 1890: 426-427, figs 11, 11a-b; With, 1908a: 249. Chelifer (Chelifer) exilimanus Balzan: Balzan, 1892: 549; Ellingsen, 1910a: 385. Neowithius exilimanus (Balzan): Beier, 1932d: 62; Beier, 1932e: 225, fig. 234; Roewer, 1937: 309; Harvey, 1991a: 649.
Type locality: Rio Apa, Paraguay. Distribution: Paraguay. Neowithius insignis (With) Chelifer insignis With, 1908a: 247-249, figs 9a-e. Neowithius insignis (With): Beier, 1932d: 62; Beier, 1932e: 224; Roewer, 1937: 310; Harvey, 1991a: 649; Muchmore, 1993a: 100.
Type locality: Contiroguita (, Venezuela?). Distribution: Venezuela? Genus Nesowithius Beier Nesowithius Beier, 1940a: 185-186; Harvey, 1991a: 649; Harvey, 1996a: 262.
Type species: Nesowithius seychellesensis Beier, 1940a, by original designation. Distribution: AFRICA: widespread; Seychelles. Nesowithius dilatimanus Mahnert Nesowithius dilatimanus Mahnert, 1988a: 43-45, figs 5-8; Harvey, 1991a: 649.
Type locality: Kilifi, Coast, Kenya. Distribution: Kenya. Nesowithius eburneus Beier Nesowithius eburneus Beier, 1979c: 112-113, fig. 6; Harvey, 1991a: 649.
Type locality: Bingerville, Abidjan, Côte d’Ivoire. Distribution: Côte d’Ivoire. Nesowithius seychellesensis Beier Nesowithius seychellesensis Beier, 1940a: 186-187, fig. 14; Mahnert, 1978d: 883-884, figs 30-32; Harvey, 1991a: 649; Judson, 1997a: 40; Harvey, 2001b: 691; Harvey, 2004: 442. Paragoniochernes digitulus Beier, 1966k: 466-467, fig. 7 (synonymised by Mahnert, 1988a: 45).
Type locality: of Nesowithius seychellesensis: Praslin Island, Seychelles. of Paragoniochernes digitulus: Durban, KwaZulu-Natal, South Africa, imported from Seychelles. Distribution: Seychelles; South Africa. Genus Parallowithius Beier Parallowithius Beier, 1955l: 318; Harvey, 1991a: 650.
Type species: Parallowithius pauper Beier, 1955l, by original designation. Distribution: AFRICA: southern Africa. Parallowithius deserticola (Beier) Allowithius deserticola Beier, 1947b: 334-335; Weidner, 1959: 113. Parallowithius deserticola (Beier): Beier, 1955l: 318; Harvey, 1991a: 650.
Type locality: Okahandja, Okahandja, Namibia. Distribution: Namibia. Parallowithius pauper Beier Parallowithius pauper Beier, 1955l: 318-320, fig. 32; Harvey, 1991a: 650.
Type locality: Cape Point, Cape Peninsula, Western Cape, South Africa. Distribution: South Africa. Genus Parawithius Chamberlin Parawithius Chamberlin, 1931c: 292; Beier, 1932e: 212; Beier, 1959e: 216; Harvey, 1991a: 650. Parawithius (Parawithius) Chamberlin: Beier, 1959e: 216; Harvey, 1991a: 650; Harvey, 2004: 437.
Type species: Chelifer nobilis With, 1908a, by original designation. Distribution: SOUTH AMERICA: Colombia; Paraguay. Parawithius iunctus Beier Chelifer (Chelifer) canestrinii Balzan: Ellingsen, 1910a: 385 (misidentification, in part). Parawithius iunctus Beier, 1932d: 57-58; Beier, 1932e: 213, fig. 219; Roewer, 1937: 309. Parawithius (Parawithius) iunctus Beier: Beier, 1959e: 216; Harvey, 1991a: 650.
Type locality: Paraguay. Distribution: Paraguay. Parawithius nobilis (With) Chelifer nobilis With, 1908a: 234-236, figs 4a-b. Parawithius nobilis (With): Chamberlin, 1931c: 292; Beier, 1932e: 213; Roewer, 1937: 309; Beier, 1959e: 216-217, fig. 25; Muchmore, 1993a: 99; Judson, 1997a: 30; Harvey, 2004: 438-439, figs 1-6; Ceballos and Florez, 2007: 50. Parawithius (Parawithius) nobilis (With): Harvey, 1991a: 650.
Type locality: Bogotá, Cundinamarca, Colombia. Distribution: Colombia. Parawithius pseudorufus Beier Chelifer (Chelifer) rufus Balzan: Ellingsen, 1910a: 386 (misidentification, in part; see Parawithius (Victorwithius) fiebrigi Beier). Parawithius pseudorufus Beier, 1932d: 58; Beier, 1932e: 214, fig. 220; Roewer, 1937: 309. Tropidowithius pseudorufus (Beier): Beier, 1955m: 11. Parawithius (Parawithius) pseudorufus Beier: Beier, 1959e: 216; Harvey, 1991a: 651.
Type locality: Paraguay. Distribution: Paraguay. Genus Plesiowithius Vachon Plesiowithius Vachon, 1954a: 1029; Harvey, 1991a: 653.
Type species: Plesiowithius dekeyseri Vachon, 1954a, by original designation. Distribution: AFRICA: western Africa. Plesiowithius dekeyseri Vachon Plesiowithius dekeyseri Vachon, 1954a: 1026-1029, figs 6-11; Harvey, 1991a: 653.
Type locality: Atar, Adrar, Mauritania. Distribution: Mauritania. Genus Pogonowithius Beier Pogonowithius Beier, 1979c: 109-111; Harvey, 1991a: 654.
Type species: Pogonowithius donisi Beier, 1979c, by original designation. Distribution: AFRICA: southern Africa. Pogonowithius donisi Beier Pogonowithius donisi Beier, 1979c: 111-112, figs 5a-b; Harvey, 1991a: 654.
Type locality: Yangambi, Haut-Zaire, Democratic Republic of Congo (Zaire). Distribution: Democratic Republic of Congo (Zaire). Genus Pycnowithius Beier Pycnowithius Beier, 1979c: 107-108; Harvey, 1991a: 654.
Type species: Pycnowithius sambicus Beier, 1979c, by original designation. Distribution: AFRICA: eastern Africa. Pycnowithius cavernicola Mahnert Pycnowithius cavernicola Mahnert, 1988a: 50-52, figs 23-28; Harvey, 1991a: 654.
Type locality: Kimakia Cave, Hunter's Lodge, Kiboko, Kenya. Distribution: Kenya. Pycnowithius garambicus (Beier) Allowithius garambicus Beier, 1972a: 13-15, fig. 7. Pycnowithius garambicus (Beier): Beier, 1979c: 108; Harvey, 1991a: 655.
Type locality: Parc National de la Garamba, Haut-Zaire, Democratic Republic of Congo (Zaire). Distribution: Democratic Republic of Congo (Zaire). Pycnowithius sambicus Beier Pycnowithius sambicus Beier, 1979c: 108-109, figs 4a-b; Harvey, 1991a: 655.
Type locality: Hangama, Zambia. Distribution: Zambia. Genus Rexwithius Heurtault Rexwithius Heurtault, 1994a: 190-191.
Type species: Rexwithius girardi Heurtault, 1994, by original designation. Distribution: AFRICA: west Africa. Rexwithius girardi Heurtault Rexwithius girardi Heurtault, 1994a: 191-198, figs 1-8.
Type locality: 10 km S. of Ndouci (as N'Douci), Abidjan, Côte d’Ivoire. Distribution: Côte d’Ivoire; Guinea. Genus Scotowithius Beier Scotowithius Beier, 1977b: 6-7; Harvey, 1991a: 655.
Type species: Scotowithius helenae Beier, 1977b, by original designation. Distribution: AFRICA: St Helena. Scotowithius helenae Beier Scotowithius helenae Beier, 1977b: 7-8, fig. 3; Harvey, 1991a: 655; Harvey, 2004: 443.
Type locality: Teutonic Hall, St Helena, St Helena. Distribution: St Helena. Genus Sphaerowithius Mahnert Sphaerowithius Mahnert, 1988a: 68-69; Harvey, 1991a: 655.
Type species: Chelifer (Trachychernes) perpusillus Ellingsen, 1910a, by original designation. Distribution: AFRICA: eastern and southern Africa. AUSTRALASIA: Solomon Islands. Sphaerowithius basilewskyi (Beier) Nannowithius basilewskyi Beier, 1962f: 30-31, fig. 14. Sphaerowithius basilewskyi (Beier): Mahnert, 1988a: 69; Harvey, 1991a: 655.
Type locality: gorge de la Mungula, Bunduki, Uluguru Mountains, Morogoro, Tanzania. Distribution: Tanzania. Sphaerowithius perpusillus (Ellingsen) Chelifer (Trachychernes) perpusillus Ellingsen, 1910a: 377-378. Chelifer perpusillus Ellingsen: Ellingsen, 1912b: 82. Nannowithius perpusillus (Ellingsen): Beier, 1932d: 57; Beier, 1932e: 211-212, fig. 218; Roewer, 1937: 309. Sphaerowithius perpusillus (Beier): Mahnert, 1988a: 45-47, figs 9-11; Harvey, 1991a: 655.
Type locality: Takaungu (as Takanuga), S. of Malindi, Coast, Kenya. Distribution: Kenya. Sphaerowithius saegeri (Beier) Nannowithius saegeri Beier, 1972a: 15-17, fig. 8. Sphaerowithius saegeri (Beier): Mahnert, 1988a: 69; Harvey, 1991a: 655.
Type locality: Parc National de la Garamba, Haut-Zaire, Democratic Republic of Congo (Zaire). Distribution: Democratic Republic of Congo (Zaire). Sphaerowithius salomonensis (Beier) Nannowithius salomonensis Beier, 1966d: 154-155, fig. 12. Sphaerowithius salomonensis (Beier): Mahnert, 1988a: 69; Harvey, 1991a: 655.
Type locality: Giro (= Gizo?), New Georgia, Western, Solomon Islands. Distribution: Solomon Islands. Sphaerowithius vafer (Beier) Nannowithius vafer Beier, 1966k: 467-469, fig. 8. Sphaerowithius vafer (Beier): Mahnert, 1988a: 69; Harvey, 1991a: 656.
Type locality: 33 miles from Pretoria via Warmbaths (as Warmbad), Limpopo, South Africa. Distribution: South Africa. Genus Sphallowithius Beier Sphallowithius Beier, 1977b: 8-9; Harvey, 1991a: 656.
Type species: Sphallowithius excelsus Beier, 1977b, by original designation. Distribution: AFRICA: St Helena. Sphallowithius excelsus Beier Sphallowithius excelsus Beier, 1977b: 9-10, fig. 4; Harvey, 1991a: 656.
Type locality: East Prosperous Bay Plain, St Helena, St Helena. Distribution: St Helena. Sphallowithius inhonestus Beier Sphallowithius inhonestus Beier, 1977b: 10-11, fig. 5; Harvey, 1991a: 656.
Type locality: Entre Lufkins and Peak Gut, St Helena, St Helena. Distribution: St Helena. Genus Stenowithius Beier Stenowithius Beier, 1932d: 55; Beier, 1932e: 206; Harvey, 1991a: 656.
Type species: Chelifer (Chelifer) buettneri Ellingsen, 1910a, by original designation. Distribution: AFRICA: widespread. Stenowithius angulatus (Ellingsen) Chelifer (Chelifer) angulatus Ellingsen, 1906: 256-258. Chelifer angulatus Ellingsen: Ellingsen, 1912b: 83. Stenowithius angulatus (Ellingsen): Beier, 1932d: 57; Beier, 1932e: 209; Roewer, 1937: 309; Harvey, 1991a: 656; Harvey, 2004: 443.
Type locality: Roça, Príncipe, São Tomé and Príncipe. Distribution: São Tomé and Príncipe. Stenowithius bayoni (Ellingsen) Chelifer bayoni Ellingsen, 1910d: 536-538; Ellingsen, 1912b: 83, 92; Ellingsen, 1913a: 453-454. Withius bayoni (Ellingsen): Godfrey, 1927: 18. Stenowithius bayoni (Ellingsen): Beier, 1932d: 57; Beier, 1932e: 207-208; Roewer, 1937: 308; Vachon, 1940f: 3; Beier, 1948b: 462; Beier, 1964k: 79; Beier, 1972a: 15; Mahnert, 1988a: 45; Harvey, 1991a: 656; Poinar, Ćurčić and Cokendolpher, 1998: 89, 90; Beron, 2002: 39; Harvey, 2004: 443. Stenowithius ugandanus Beier, 1932d: 55-56; Beier, 1932e: 207, fig. 213; Roewer, 1937: 308; Beier, 1948b: 445; Beier, 1955c: 556; Judson, 1997a: 44 (as junior synonym of Stenowithius bayoni (Ellingsen)) (synonymised by Beier, 1964k: 79). Not Stenowithius ugandanus Beier: Caporiacco, 1949a: 318 (misidentification; see Withius congicus (Beier)).
Type locality: of Chelifer bayoni: Buvama, Uganda; and Bugala, Uganda. of Stenowithius ugandanus: Kampala, Uganda. Distribution: Democratic Republic of Congo (Zaire); Kenya; Malawi; Mozambique; South Africa; Uganda. Stenowithius bayoni angustus Beier Stenowithius bayoni angustus Beier, 1964k: 79-80, fig. 34; Harvey, 1991a: 656.
Type locality: Potgietersrus (as Potgietersrust), Limpopo, South Africa. Distribution: South Africa. Stenowithius bayoni bayoni (Ellingsen)
Distribution: Democratic Republic of Congo (Zaire); Kenya; Malawi; Mozambique; South Africa; Uganda. Stenowithius buettneri (Ellingsen) Chelifer (Chelifer) buettneri Ellingsen, 1910a: 383-384 (Chelifer (Chelifer) büttneri). Chelifer buettneri Ellingsen: Ellingsen, 1912b: 83 (as Chelifer büttneri) Stenowithius buettneri (Ellingsen): Beier, 1932d: 55 (as Stenowithius büttneri); Beier, 1932e: 209, fig. 215 (as Stenowithius büttneri); Roewer, 1937: 309 (as Stenowithius büttneri); Mahnert, 1988a: 66, figs 54-56; Harvey, 1991a: 657.
Type locality: Bismarckbourg, Centrale, Togo. Distribution: Malawi; Togo. Stenowithius duffeyi Beier Stenowithius duffeyi Beier, 1961c: 597-598, fig. 3; Duffey, 1964: 241, 250, fig. 6; Harvey, 1991a: 657; Ashmole and Ashmole, 1997: 566, 579, fig. 8; Judson, 1997a: 15; Harvey, 2004: 443.
Type locality: Boatswainbird (as Bos'nbird) Island, Ascension Island, St Helena. Distribution: St Helena (Ascension Island). Stenowithius parvulus Beier Stenowithius parvulus Beier, 1954d: 136-137, fig. 4; Harvey, 1991a: 657.
Type locality: Flandria, Equateur, Democratic Republic of Congo (Zaire). Distribution: Democratic Republic of Congo (Zaire). Stenowithius persimilis Beier Stenowithius persimilis Beier, 1932d: 56-57; Beier, 1932e: 208, fig. 214; Roewer, 1937: 309; Beier, 1954d: 136; Harvey, 1991a: 657.
Type locality: Rutshuru (as Rutshum [sic]), Buseregenye (as Buseregenye), Nord-Kivu, Democratic Republic of Congo (Zaire). Distribution: Democratic Republic of Congo (Zaire). Stenowithius phagophilus Beier Stenowithius phagophilus Beier, 1953c: 78, fig. 3; Harvey, 1991a: 657.
Type locality: Aminuis Reserve, Hereroland Oss, Namibia. Distribution: Namibia. Stenowithius torpidus Beier Stenowithius torpidus Beier, 1958a: 181-183, fig. 13; Harvey, 1991a: 657.
Type locality: Pietermaritzburg, KwaZulu-Natal, South Africa. Distribution: South Africa. Genus Termitowithius Muchmore Termitowithius Muchmore, 1990b: 125.
Type species: Termitowithius kistneri Muchmore, 1990b, by original designation. Distribution: AFRICA: Tanzania. Termitowithius kistneri Muchmore Termitowithius kistneri Muchmore, 1990b: 126-127, figs 1-7.
Type locality: Lake Manyara National Park, Arusha, Tanzania. Distribution: Tanzania. Genus Thaumatowithius Beier Thaumatowithius Beier, 1940a: 187-188; Harvey, 1991a: 657.
Type species: Thaumatowithius tibialis Beier, 1940a, by original designation. Distribution: AFRICA: Mascarene Islands. Thaumatowithius aberrans Mahnert Thaumatowithius aberrans Mahnert, 1975b: 549-553, figs 4a-k, 6f; Harvey, 1991a: 657.
Type locality: Plaine des Chicots, Réunion. Distribution: Réunion. Thaumatowithius tibialis Beier Thaumatowithius tibialis Beier, 1940a: 188-189, fig. 15; Harvey, 1991a: 657; Judson, 1997a: 43; Harvey, 2004: 443.
Type locality: Le Pouce, Mauritius, Mauritius. Distribution: Mauritius. Genus Trichotowithius Beier Trichotowithius Beier, 1944: 203-204; Harvey, 1991a: 658; Dashdamirov, 1992: 297-298.
Type species: Trichotowithius abyssinicus Beier, 1944, by original designation. Distribution: AFRICA: eastern Africa. Trichotowithius abyssinicus Beier Trichotowithius abyssinicus Beier, 1944: 204-206, fig. 20; Harvey, 1991a: 658; Dashdamirov, 1992: 294-297, figs 1-15; Judson, 1997a: 7; Beron, 2002: 39; Harvey, 2004: 443. Trichotowithius elgonensis Beier, 1955c: 556-557, fig. 19; Mahnert, 1988a: 48; Harvey, 1991a: 658; Beron, 2002: 39 (synonymised by Dashdamirov, 1992: 294).
Type locality: of Trichotowithius abyssinicus: Jemme Forest (as Djem-Djem Wald), Ethiopia. of Trichotowithius elgonensis: Mt Elgon, E. side, Western, Kenya. Distribution: Ethiopia; Kenya. Genus Tropidowithius Beier Tropidowithius Beier, 1955m: 10-11; Harvey, 1991a: 658.
Type species: Tropidowithius peruanus Beier, 1955m, by original designation. Distribution: SOUTH AMERICA: Peru. Tropidowithius peruanus Beier Tropidowithius peruanus Beier, 1955m: 11-12, fig. 11; Weidner, 1959: 117; Harvey, 1991a: 658; Mahnert and Adis, 2002: 380.
Type locality: Sivia, Ayacucho, Peru. Distribution: Peru. Genus Victorwithius Feio Victorwithius Feio, 1944: 1-3; Harvey, 2004: 439-440. Cacodemoniellus Beier, 1954c: 326-327 (synonymised by Beier, 1959e: 216). Parawithius (Victorwithius) Feio: Beier, 1959e: 216; Harvey, 1991a: 651.
Type species: of Victorwithius: Victorwithius monoplacophorus Feio, 1944, by original designation. of Cacodemoniellus: Cacodemoniellus mimulus Beier, 1954c, by original designation. Distribution: SOUTH AMERICA: widespread. Victorwithius coniger (Mahnert) Parawithius (Victorwithius) coniger Mahnert, 1979d: 804-806, figs 182-186; Mahnert and Adis, 1986: 214; Adis, Junk and Penny, 1987: 489; Harvey, 1991a: 651; Mahnert and Adis, 2002: 380. Victorwithius coniger (Mahnert): Harvey, 2004: 440, 443.
Type locality: Rio Solimões, Manaus, Amazonas, Brazil. Distribution: Brazil (Amazonas). Victorwithius fiebrigi (Beier) Chelifer (Chelifer) rufus Balzan: Ellingsen, 1910a: 386 (misidentification, in part; see Parawithius (Parawithius) pseudorufus Beier). Parawithius fiebrigi Beier, 1932d: 59; Beier, 1932e: 215, fig. 222; Roewer, 1937: 309; Feio, 1945: 7. Tropidowithius fiebrigi (Beier): Beier, 1955m: 11. Parawithius (Victorwithius) fiebrigi Beier: Beier, 1959e: 216; Harvey, 1991a: 651. Victorwithius fiebrigi (Beier): Harvey, 2004: 440.
Type locality: Paraguay. Distribution: Argentina; Paraguay. Victorwithius gracilimanus (Mahnert) Parawithius (Victorwithius) gracilimanus Mahnert, 1979d: 802-804, figs 176-181; Adis, 1981: 119, etc.; Adis and Mahnert, 1986: 312; Mahnert and Adis, 1986: 214; Adis, Junk and Penny, 1987: 489; Mahnert, Adis, and Bührnheim, 1986: fig. 16; Harvey, 1991a: 651; Aguiar and Bührnheim, 1992c: 199-200, figs 2-5; Aguiar and Bührnheim, 1998a: 387; Aguiar and Bührnheim, 1998b: 456; Mahnert and Adis, 2002: fig. 14. Parawithius gracilimanus Mahnert: Aguiar and Bührnheim, 1994: 74; Poinar, Ćurčić and Cokendolpher, 1998: 86, 87; Mahnert and Adis, 2002: 380. Victorwithius gracilimanus (Mahnert): Harvey, 2004: 440.
Type locality: Río Tarumã-Mirím, Manaus, Amazonas, Brazil. Distribution: Brazil (Acre, Amazonas, Roraima). Victorwithius incognitus (Beier) Parawithius (Victorwithius) incognitus Beier, 1959e: 220-221, fig. 29; Harvey, 1991a: 651; Mahnert and Adis, 2002: 380. Victorwithius incognitus (Beier): Harvey, 2004: 440.
Type locality: 18 miles NE. of La Merced, Río Perené, Colonia Perené, Junín, Peru. Distribution: Peru. Victorwithius mimulus (Beier) Cacodemoniellus mimulus Beier, 1954c: 327-329, fig. 2. Parawithius (Victorwithius) mimulus (Beier): Beier, 1959e: 216; Harvey, 1991a: 652. Victorwithius mimulus (Beier): Harvey, 2004: 440.
Type locality: Palmeira, Paraná, Brazil. Distribution: Brazil (Paraná). Victorwithius monoplacophorus Feio Victorwithius monoplacophorus Feio, 1944: 1-3, figs 1-11; Feio, 1945: 8; Kury and Nogueira, 1999: 10; Harvey, 2004: 440. Parawithius (Victorwithius) monoplacophorus (Feio): Beier, 1959e: 216; Harvey, 1991a: 652.
Type locality: Etienne, Bituruna, Paraná, Brazil. Distribution: Argentina; Brazil (Paraná); Uruguay. Victorwithius proximus (Ellingsen) Chelifer (Chelifer) proximus Ellingsen, 1905e: 324-326; With, 1908a: 231. Parawithius proximus (Ellingsen): Chamberlin, 1931c: 292; Beier, 1932e: 215; Roewer, 1937: 309; Mello-Leitão, 1939a: 122; Mello-Leitão, 1939b: 617. Parawithius (Victorwithius) proximus (Ellingsen): Beier, 1959e: 216; Harvey, 1991a: 652. Victorwithius proximus (Ellingsen): Harvey, 2004: 440.
Type locality: Santa Catalina, Jujuy, Argentina. Distribution: Argentina. Victorwithius rufeolus (Beier) Parawithius (Victorwithius) rufeolus Beier, 1959e: 221-222, fig. 30; Muchmore, 1977: 77; Harvey, 1991a: 652; Muchmore, 1991b: 158; Ceballos, 2004: 42 |