Pseudoscorpions of the World

Last updated: 19 January 2009

Version 1.2, 2009

Text Box: Atemnidae	Text Catalogue
Bochicidae	Text Catalogue
Cheiridiidae	Text Catalogue
Cheliferidae	Text Catalogue
Chernetidae	Text Catalogue
Chthoniidae	Text Catalogue
Dracochelidae	Text Catalogue
Feaellidae	Text Catalogue
Garypidae	Text Catalogue
Garypinidae	Text Catalogue
Geogarypidae	Text Catalogue
Gymnobisiidae	Text Catalogue
Hyidae	Text Catalogue
Ideoroncidae	Text Catalogue
Larcidae	Text Catalogue
Lechytiidae	Text Catalogue
Menthidae	Text Catalogue
Neobisiidae	Text Catalogue
Olpiidae	Text Catalogue
Parahyidae	Text Catalogue
Pseudochiridiidae	Text Catalogue
Pseudogarypidae	Text Catalogue
Sternophoridae 	Text Catalogue
Syarinidae 	Text Catalogue
Tridenchthoniidae	Text Catalogue
Withiidae	Text Catalogue
Text Box: Home
Text Box: What are pseudoscorpions?
Text Box: Phylogeny
Text Box: Classification & list of families
Text Box: Literature
Text Box: List of genera, alphabetical
Text Box: Numbers of Recent taxa 
Text Box: Fossils
Text Box: Full classification 
Text Box: Nomina dubia & nomina nuda
Text Box: Species by country

ORDER PSEUDOSCORPIONES DE GEER

 

Suborder Epiocheirata Harvey 1992

Superfamily Chthonioidea Daday 1888

                          Family Chthoniidae Daday 1888

                                       Subfamily Chthoniinae Daday 1888

                                                    Tribe Chthoniini Daday 1888

Anisoditha Chamberlin and R.V. Chamberlin 1945

Aphrastochthonius Chamberlin 1962

Apochthonius Chamberlin 1929

Austrochthonius Chamberlin 1929

Caribchthonius Muchmore 1976

Chiliochthonius Vitali-di Castri 1976

Chthonius C.L. Koch 1843

Congochthonius Beier 1959

Drepanochthonius Beier 1964

Francochthonius Vitali-di Castri 1976

Kleptochthonius Chamberlin 1949

Malcolmochthonius Benedict 1978

Mexichthonius Muchmore 1975

Mundochthonius Chamberlin 1929

Neochthonius Chamberlin 1929

Pseudochthonius Balzan 1892

Sathrochthoniella Beier 1967

Sathrochthonius Chamberlin 1962

Spelyngochthonius Beier 1955

Tyrannochthoniella Beier 1966

                                                    Tribe Tyrannochthoniini Chamberlin 1962

Lagynochthonius Beier 1951

Maorichthonius Chamberlin 1925

Paraliochthonius Beier 1956

Troglochthonius Beier 1939

Tyrannochthonius Chamberlin 1929

Vulcanochthonius Muchmore 2000

                                                    Subfamily Pseudotyrannochthoniinae Beier 1930

Afrochthonius Beier 1930

Allochthonius Chamberlin 1929

Centrochthonius Beier 1931

Pseudotyrannochthonius Beier 1930

Selachochthonius Chamberlin 1929

 

† Family Dracochelidae Schawaller, Shear and Bonamo 1990

Dracochela Schawaller, Shear and Bonamo 1990

 

         Family Lechytiidae Chamberlin 1929

Lechytia Balzan 1892

                         

                                       Family Tridenchthoniidae Balzan 1892

                                                    Subfamily Tridenchthoniinae Balzan 1892

Chelignathus Menge 1854

Compsaditha Chamberlin 1929

Ditha Chamberlin 1929

Dithella Chamberlin and R.V. Chamberlin 1945

Haploditha Caporiacco 1951

Heterolophus Tömösváry 1884

Neoditha Feio 1945

Tridenchthonius Balzan 1887

Typhloditha Beier 1955

                                                    Subfamily Verrucadithinae Chamberlin 1929

Anaulacodithella Beier 1944

Cryptoditha Chamberlin and R.V. Chamberlin 1945

Pycnodithella Beier 1947

Rheodithella Dashdamirov and Judson 2004

Sororoditha Chamberlin and R.V. Chamberlin 1945

Verrucaditha Chamberlin 1929

Verrucadithella Beier 1931

 

Superfamily Feaelliodea Ellingsen 1906

         Family Feaellidae Ellingsen 1906

Feaella Ellingsen 1906

 

                                       Family Pseudogarypidae Chamberlin 1923

Neopseudogarypus Morris 1948

Pseudogarypus Ellingsen 1909

 

Suborder Iocheirata Harvey 1992

                          Superfamily Neobisioidea Chamberlin 1930

                                       Family Bochicidae Chamberlin 1930

                                                    Subfamily Bochicinae Chamberlin 1930

Antillobisium Dumitresco and Orghidan 1977

Bochica Chamberlin 1930

Spelaeobochica Mahnert 2001

Troglobisium Beier 1939

Troglobochica Muchmore 1984

Troglohya Beier 1956

Vachonium Chamberlin 1947

                                                    Subfamily Leucohyinae Chamberlin 1946

Apohya Muchmore 1973

Leucohya Chamberlin 1946

Mexobisium Muchmore 1972

Paravachonium Beier 1956

            

         Family Gymnobisiidae Beier 1947

Beierobisium Vitali-di Castri 1970

Gymnobisium Beier 1931

Mirobisium Beier 1931

Vachonobisium Vitali-di Castri 1963

 

                                       Family Hyidae Chamberlin 1930

                                                    Subfamily Hyinae Chamberlin 1930

Hya Chamberlin 1930

                                                    Subfamily Indohyinae Harvey 1993

Indohya Beier 1974

                                      

         Family Ideoroncidae Chamberlin 1930

Afroroncus Mahnert 1981

Albiorix Chamberlin 1930

Dhanus Chamberlin 1930

Ideoroncus Balzan 1887

Nannoroncus Beier 1955

Negroroncus Beier 1931

Nhatrangia Redikorzev 1938

Pseudalbiorix Harvey, Barba, Muchmore and Pérez 2007

Shravana Chamberlin 1930

Typhloroncus Muchmore 1979

Xorilbia Harvey and Mahnert 2006

                                      

                                       Family Neobisiidae Chamberlin 1930

                                                    Subfamily Microcreagrinae Balzan 1892

Acanthocreagris Mahnert 1974

Alabamocreagris Ćurčić 1984

Americocreagris Ćurčić 1982

Australinocreagris Ćurčić 1984

Bisetocreagris Ćurčić 1983

Cryptocreagris Ćurčić 1984

Dentocreagris Dashdamirov 1997

Fissilicreagris Ćurčić 1984

Globocreagris Ćurčić 1984

Halobisium Chamberlin 1930

Insulocreagris Ćurčić 1987

Lissocreagris Ćurčić 1981

Microcreagris Balzan 1892

Minicreagris Ćurčić 1989

Orientocreagris Ćurčić 1985

Roncocreagris Mahnert 1974

Saetigerocreagris Ćurčić 1984

Stenohya Beier 1967

Tartarocreagris Ćurčić 1984

Tuberocreagris Ćurčić 1978

                                                    Subfamily Neobisiinae Chamberlin 1930

Balkanoroncus Ćurčić 1975

Ernstmayria Ćurčić and Dimitrijević 2006

Microbisium Chamberlin 1930

Neobisium Chamberlin 1930

Novobisium Muchmore 1967

Occitanobisium Heurtault 1977

Paedobisium Beier 1939

Parobisium Chamberlin 1930

Protoneobisium Ćurčić 1988

Roncobisium Vachon 1967

Roncus L. Koch 1873

Trisetobisium Ćurčić 1982

            

                                       Family Parahyidae Harvey 1992

Parahya Beier 1957

 

                                       Family Syarinidae Chamberlin 1930

                                                    Subfamily Chitrellinae Beier 1932

Aglaochitra Chamberlin 1952

Chitrella Beier 1932

Chitrellina Muchmore 1996

Hadoblothrus Beier 1952

Microcreagrella Beier 1961

Microcreagrina Beier 1961

Pararoncus Chamberlin 1938

Pseudoblothrus Beier 1931

                                                    Subfamily Ideobisiinae Banks 1895

Alocobisium Beier 1952

Ideobisium Balzan 1892

Ideoblothrus Balzan 1892

Microblothrus Mahnert 1985

Nannobisium Beier 1931

                                                    Subfamily Syarininae Chamberlin 1930

Anysrius Harvey 1998

Syarinus Chamberlin 1925

                                                    Unplaced

Hyarinus Chamberlin 1925

 

                          Superfamily Garypoidea Simon 1879

         Family Garypidae Simon 1879

Ammogarypus Beier 1962

Anagarypus Chamberlin 1930

Elattogarypus Beier 1964

Eremogarypus Beier 1955

Garypus L. Koch 1873

Meiogarypus Beier 1955

Neogarypus Vachon 1937

Paragarypus Vachon 1937

Synsphyronus Chamberlin 1930

Thaumastogarypus Beier 1947

 

                                       Family Garypinidae Daday 1888

Aldabrinus Chamberlin 1930

Amblyolpium Simon 1898

Caecogarypinus Dashdamirov 2007

Galapagodinus Beier 1978

Garypinidius Beier 1955

Garypinus Daday 1888

Haplogarypinus Beier 1959

Hemisolinus Beier 1977

Indogarypinus Murthy and Ananthakrishnan 1977

Nelsoninus Beier 1967

Neoamblyolpium Hoff 1956

Neominniza Beier 1930

Oreolpium Benedict and Malcolm 1978

Paraldabrinus Beier 1966

Protogarypinus Beier 1954

Pseudogarypinus Beier 1931

Serianus Chamberlin 1930

Solinellus Muchmore 1979

Solinus Chamberlin 1930

Teratolpium Beier 1959

Thaumatolpium Beier 1931

 

                                       Family Geogarypidae Chamberlin 1930

Afrogarypus Beier 1931

Geogarypus Chamberlin 1930

Indogarypus Beier 1957

 

                                       Family Larcidae Harvey 1992

Archeolarca Hoff and Clawson 1952

Larca Chamberlin 1930

 

                                       Family Menthidae Chamberlin 1930

Menthus Chamberlin 1930

Oligomenthus Beier 1962

Paramenthus Beier 1963

Pseudomenthus Mahnert 2007

Thenmus Harvey 1990

 

                                       Family Olpiidae Banks 1895

                                                                 Tribe Hesperolpiini Hoff 1964

Aphelolpium Hoff 1964

Apolpium Chamberlin 1930

Calocheirus Chamberlin 1930

Cardiolpium Mahnert 1986

Ectactolpium Beier 1947

Hesperolpium Chamberlin 1930

Nanolpium Beier 1947

Planctolpium Hoff 1964

Progarypus Beier 1931

Stenolpiodes Beier 1959

Stenolpium Beier 1955

                                                                 Tribe Olpiini Bansk 1895

Antillolpium Muchmore 1991

Austrohorus Beier 1966

Banksolpium Muchmore 1986

Beierolpium Heurtault 1976

Calocheiridius Beier and Turk 1952

Euryolpium Redikorzev 1938

Halominniza Mahnert 1975

Heterolpium Sivaraman 1980

Hoffhorus Heurtault 1976

Horus Chamberlin 1930

Indolpium Hoff 1945

Leptolpium Tooren 2002

Minniza Simon 1881

Neopachyolpium Hoff 1945

Nipponogarypus Morikawa 1955

Novohorus Hoff 1945

Olpiolum Beier 1931

Olpium L. Koch 1873

Pachyolpium Beier 1931

Parolpium Beier 1931

Pseudohorus Beier 1946

Xenolpium Chamberlin 1930

 

                          Superfamily Sternophoroidea Chamberlin 1923

                                       Family Sternophoridae Chamberlin 1923

Afrosternophorus Beier 1967

Garyops Banks 1909

Idiogaryops Hoff 1963

 

                          Superfamily Cheiridioidea Hansen 1894

                                       Family Cheiridiidae Hansen 1894

                                                    Subfamily Cheiridiinae Hansen 1894

Apocheiridium Chamberlin 1924

Cheiridium Menge 1855

Cryptocheiridium Chamberlin 1931

Neocheiridium Beier 1932

Nesocheiridium Beier 1957

            Subfamily Pycnocheiridiinae Hansen 1894

Pycnocheiridium Beier 1964

 

         Family Pseudochiridiidae Chamberlin 1923

Paracheiridium Vachon 1938

Pseudochiridium With 1906

 

                          Superfamily Cheliferoidea Risso 1826

                                       Family Atemnidae Chamberlin 1931

                                                    Subfamily Atemninae Chamberlin 1931

Anatemnus Beier 1932

Atemnus Canestrini 1884

Athleticatemnus Beier 1979

Catatemnus Beier 1932

Cyclatemnus Beier 1932

Mesatemnus Beier and Turk 1952

Metatemnus Beier 1932

Micratemnus Beier 1932

Oratemnus Beier 1932

Paratemnoides Harvey 1991

Progonatemnus Beier 1955

Stenatemnus Beier 1932

Synatemnus Beier 1944

Tamenus Beier 1932

Titanatemnus Beier 1932

                                                    Subfamily Miratemninae Beier 1932

Brazilatemnus Muchmore 1975

Caecatemnus Mahnert 1985

Diplotemnus Chamberlin 1933

Miratemnus Beier 1932

Tullgrenius Chamberlin 1933

            

                                       Family Cheliferidae Risso 1826

                      Subfamily Cheliferinae Risso 1826

Amaurochelifer Beier 1951

Ancistrochelifer Beier 1951

Aperittochelifer Beier 1955

Aporochelifer Beier 1953

Aspurochelifer Benedict and Malcolm 1979

Australochelifer Beier 1975

Beierius Chamberlin 1932

Beierochelifer Mahnert 1977

Canarichelifer Beier 1965

Centrochelifer Beier 1959

Chamberlinarius Heurtault 1990

Cheirochelifer Beier 1967

Chelifer Geoffroy 1762

Cubachelifer Hoff 1946

Dactylochelifer Beier 1932

Dichela Menge 1854

Electrochelifer Beier 1937

Ellingsenius Chamberlin 1932

Eremochernes Beier 1932

Florichelifer Hoff 1964

Gobichelifer Krumpál 1979

Hansenius Chamberlin 1932

Haplochelifer Chamberlin 1932

Hygrochelifer Murthy and Ananthakrishnan 1977

Hysterochelifer Chamberlin 1932

Idiochelifer Chamberlin 1932

Kashimachelifer Morikawa 1957

Levichelifer Hoff 1946

Lissochelifer Chamberlin 1932

Litochelifer Beier 1948

Lophochernes Simon 1878

Lophodactylus Chamberlin 1932

Macrochelifer Vachon 1940

Mesochelifer Vachon 1940

Metachelifer Redikorzev 1938

Mexichelifer Muchmore 1973

Microchelifer Beier 1944

Mucrochelifer Beier 1932

Nannochelifer Beier 1967

Nannocheliferoides Beier 1974

Pachychelifer Beier 1962

Paisochelifer Hoff 1946

Papuchelifer Beier 1965

Parachelifer Chamberlin 1932

Phorochelifer Hoff 1956

Pilochelifer Beier 1935

Protochelifer Beier 1948

Pseudorhacochelifer Beier 1976

Pugnochelifer Hoff 1964

Pycnochelifer Beier 1937

Rhacochelifer Beier 1932

Rhopalochelifer Beier 1964

Sinochelifer Beier 1967

Stenochelifer Beier 1967

Strobilochelifer Beier 1932

Stygiochelifer Beier 1932

Telechelifer Chamberlin 1949

Tetrachelifer Beier 1967

Trachychelifer Hong, 1983

Tyrannochelifer Chamberlin 1932

Xenochelifer Chamberlin 1949

                                                    Subfamily Philomaoriinae Chamberlin 1931

Philomaoria Chamberlin 1931

                                                    Unplaced

Ectoceras Stecker 1875

 

                                       Family Chernetidae Menge 1855

                      Subfamily Chernetinae Menge 1855

                                   Tribe Chernetini Menge 1855

Acanthicochernes Beier 1964

Acuminochernes Hoff 1949

Adelphochernes Beier 1937

Americhernes Muchmore 1976

Anaperochernes Beier 1964

Antillochernes Muchmore 1984

Apatochernes Beier 1948

Asterochernes Beier 1955

Atherochernes Beier 1954

Austrochernes Beier 1932

Barbaraella Harvey 1995

Bituberochernes Muchmore 1974

Byrsochernes Beier 1959

Cacoxylus Beier 1965

Caffrowithius Beier 1932

Calidiochernes Beier 1954

Caribochernes Beier 1976

Ceratochernes Mahnert 1994

Ceriochernes Beier 1937

Chelanops Gervais 1849

Chelodamus R.V. Chamberlin 1925

Chernes Menge 1855

Chiridiochernes Muchmore 1972

Chrysochernes Hoff 1956

Cocinachernes Hentschel and Muchmore 1989

Coprochernes Beier 1976

Cordylochernes Beier 1932

Corosoma Karsch 1879

Cyclochernes Beier 1970

Dasychernes Chamberlin 1929

Dendrochernes Beier 1932

Dinocheirus Chamberlin 1929

Dinochernes Beier 1933

Diplothrixochernes Beier 1962

Epactiochernes Muchmore 1974

Epichernes Muchmore 1982

Eumecochernes Beier 1932

Gelachernes Beier 1940

Gigantochernes Beier 1932

Gomphochernes Beier 1932

Haplochernes Beier 1932

Hebridochernes Beier 1940

Hesperochernes Chamberlin 1924

Heterochernes Beier 1966

Hexachernes Beier 1953

Illinichernes Hoff 1949

Incachernes Beier 1933

Indochernes Murthy and Ananthakrishnan 1977

Interchernes Muchmore 1980

Lustrochernes Beier 1932

Macrochernes Hoff 1946

Maorichernes Beier 1932

Maxchernes Feio 1960

Meiochernes Beier 1957

Mesochernes Beier 1932

Metagoniochernes Vachon 1939

Mexachernes Hoff 1947

Mirochernes Beier 1930

Mucrochernes Muchmore 1973

Neoallochernes Hoff 1947

Neochelanops Beier 1964

Neochernes Beier 1932

Nesidiochernes Beier 1957

Nesiotochernes Beier 1976

Nesochernes Beier 1932

Ochrochernes Beier 1932

Odontochernes Beier 1932

Oligochernes Beier 1937

Opsochernes Beier 1966

Orochernes Beier 1968

Pachychernes Beier 1932

Paracanthicochernes Beier 1966

Parachernes Chamberlin 1931

Parapilanus Beier 1973

Paraustrochernes Beier 1966

Parazaona Beier 1932

Petterchernes Heurtault 1986

Phaulochernes Beier 1976

Phymatochernes Mahnert 1979

Pilanus Beier 1930

Pseudopilanus Beier 1957

Reischekia Beier 1948

Rhinochernes Beier 1955

Rhopalochernes Beier 1932

Satapranus Harvey and Volschenk 2007

Semeiochernes Beier 1932

Smeringochernes Beier 1957

Spelaeochernes Mahnert 2001

Sphenochernes Turk 1953

Sundochernes Beier 1932

Sundowithius Beier 1932

Systellochernes Beier 1964

Teratochernes Beier 1957

Thalassochernes Beier 1940

Thapsinochernes Beier 1957

Troglochernes Beier 1969

Tuberochernes Muchmore 1997

Tychochernes Hoff 1956

Verrucachernes Chamberlin 1947

Wyochernes Hoff 1949

Zaona Chamberlin 1925

                                                                 Tribe Myrmochernetini Chamberlin 1931

Marachernes Harvey 1992

Myrmochernes Tullgren 1907

Xenochernes Feio 1945

                                                    Subfamily Goniochernetinae Beier 1932

Calymmachernes Beier 1954

Conicochernes Beier 1948

Goniochernes Beier 1932

            Subfamily Lamprochernetinae Beier 1932

Allochernes Beier 1932

Anthrenochernes Lohmander 1939

Bipeltochernes Dashdamirov 2005

Lamprochernes Tömösváry 1882

Lasiochernes Beier 1932

Megachernes Beier 1932

Nudochernes Beier 1935

Pselaphochernes Beier 1932

                                      

         Family Withiidae Chamberlin 1931

                      Subfamily Paragoniochernetinae Beier 1944

Cyrtowithius Beier 1955

Ectromachernes Beier 1944

Paragoniochernes Beier 1932

Pseudatemnus Beier 1947

Pseudochernes Beier 1954

                                                    Subfamily Withiinae Chamberlin 1931

                                                                 Tribe Cacodemoniini Chamberlin 1931

Cacodemonius Chamberlin 1931

                                                                 Tribe Juxtacheliferini Hoff 1956

Juxtachelifer Hoff 1956

                                                                 Tribe Protowithiini Beier 1955

Protowithius Beier 1955

                                                                 Tribe Withiini Chamberlin 1931

Afrowithius Chamberlin 1931

Aisthetowithius Beier 1967

Balanowithius Beier 1959

Beierowithius Mahnert 1979

Cryptowithius Beier 1967

Cystowithius Harvey 2004

Dolichowithius Chamberlin 1931

Girardwithius Heurtault 1994

Hyperwithius Beier 1951

Metawithius Chamberlin 1931

Nannowithius Beier 1932

Neowithius Beier 1932

Nesowithius Beier 1940

Parallowithius Beier 1955

Parawithius Chamberlin 1931

Plesiowithius Vachon 1954

Pogonowithius Beier 1979

Pycnowithius Beier 1979

Rexwithius Heurtault 1994

Scotowithius Beier 1977

Sphaerowithius Mahnert 1988

Sphallowithius Beier 1977

Stenowithius Beier 1932

Termitowithius Muchmore 1990

Thaumatowithius Beier 1940

Trichotowithius Beier 1944

Tropidowithius Beier 1955

Victorwithius Feio 1944

Withius Kew 1911

 

Classification, list of valid genera