Pseudoscorpions of the World

Last updated: 19 January 2009

Version 1.2, 2009

Subfamily NEOBISIINAE Chamberlin

Neobisiinae Chamberlin, 1930: 9; Chamberlin, 1931a: 217; Beier, 1932a: 77; Roewer, 1937: 244; Hoff, 1949b: 444; Hoff, 1956b: 2; Morikawa, 1960: 112.

Genus Balkanoroncus Ćurčić

Balkanoroncus Ćurčić, 1975b: 144; Ćurčić, 1978b: 119, 131; Harvey, 1991a: 330.

 

Type species: Roncus (Parablothrus) bureschi Hadži, 1939b (now Balkanoroncus hadzii Harvey, 1991), by original designation.

Distribution: EUROPE: southern Europe.

Balkanoroncus boldorii (Beier)

Neobisium (Blothrus) boldorii Beier, 1931c: 12-13, fig. 4; Beier, 1932a: 110, fig. 129; Roewer, 1937: 248.

Neobisium boldorii Beier: Boldori, 1932: 127; Wolf, 1938: 609.

Roncus (Parablothrus) boldorii (Beier): Beier, 1942: 134; Beier, 1958b: 163; Vachon and Gabbutt, 1964, fig. 14; Beier, 1970f: 320; Gardini, 1980c: 112, 123, 126; Mahnert, 1980g: 98-99, figs 4-7.

Roncus (Parablothrus) ghidinii Beier, 1942: 134-136, fig. 3; Beier, 1972c: 3-5, fig. 1; Costantini, 1976: 123; Gardini, 1980c: 113, 123, 126 (synonymised by Mahnert, 1980h: 98).

Roncus (Parablothrus) boldorii boldorii (Beier): Beier, 1963b: 201, fig. 212; Lazzeroni, 1969b: 407; Costantini, 1976: 123.

Roncus (Parablothrus) boldorii ghidinii Beier: Beier, 1963b: 201.

Balkanoroncus baldensis Gardini, 1982d: 161-169, figs 1-15; Gardini, 1985b: 61 (synonymised by Gardini and Rizzerio, 1986a: 49).

Balkanoroncus boldorii (Beier): Gardini and Rizzerio, 1986a: 49-51; Sciaky and Vigna Taglianti, 1990: 170; Gardini, 1991a: 200-201, fig. 14; Harvey, 1991a: 330-331; Caoduro, Osella and Ruffo, 1994: 36-37; Gardini, 1995a: 5; Gardini, 1996: 161; Gardini, 2000: 102-103.

 

Type locality: of Neobisium (Blothrus) boldorii: Büs Coalghés, Gavardo, Lombardia, Italy.

of Roncus (Parablothrus) ghidinii: Büs Soradùr (as Bus Suradur), Lombardia, Italy.

of Balkanoroncus baldensis: Grotte dei Cervi, Monte Baldo, Trentino-Alto Adige, Italy.

Distribution: Italy (mainland).

Balkanoroncus bureschi (Redikorzev)

Obisium (Blothrus) bureschi Redikorzev, 1928: 129-133, figs 8-9, 10c.

Roncus (Parablothrus) bureschi (Redikorzev): Beier, 1928: 309.

Neobisium (Blothrus) bureschi (Redikorzev): Beier, 1932a: 113, fig. 134; Roewer, 1937: 248; Beier, 1939d: 72.

Neobisium bureschi (Redikorzev): Wolf, 1938: 610.

Neobisium (Heoblothrus) bureschi (Redikorzev): Beier, 1963b: 134-135, fig. 134; Beier, 1970f: 320.

Balkanoroncus praeceps Ćurčić, 1978b: 125-129, figs 7-20 (invalid replacement name for Obisium (Blothrus) bureschi Redikorzev).

Balkanoroncus bureschi (Redikorzev): Harvey, 1991a: 331.

Balkanoroncus hadzii Harvey: Petrov, 1997: 264 (lapsus).

 

Type locality: Cave Seevata dupka (as Seewa Dupka Cave), near Malka Bresnitza, S. of Teteven (as Tetewen), Lovech, Bulgaria.

Distribution: Bulgaria.

Balkanoroncus hadzii Harvey

Roncus (Parablothrus) bureschi Hadži, 1939b: 34-37, figs 9a-m (junior secondary homonym of Balkanoroncus bureschi (Redikorzev)).

Microcreagris bureschi (Hadži): Beier, 1963b: 219; Beier, 1970f: 321.

Acanthocreagris (?) bureschi (Hadži): Mahnert, 1974d: 849.

Balkanoroncus bureschi (Hadži): Ćurčić, 1975b: 144; Ćurčić, 1978b: 120-125, figs 1-6, 6a-c.

Balkanoroncus hadzii Harvey, 1991a: 331; Petrov, 1997: 264.

 

Type locality: Cave Razishkata peshtera (as Gornjata Peschtera Cave), Lakatnik, Iskur River gorge, Sofiya, Bulgaria.

Distribution: Bulgaria.

Genus Ernstmayria Ćurčić and Dimitrijević

Ernstmayria Ćurčić and Dimitrijević, in Ćurčić, Dimitrijević, Legakis, Makarov, Tomić, S.B. Ćurčić, Mitić and Lučić, 2006: 39.

 

Type species: Ernstmayria apostolostrichasi Ćurčić, Dimitrijević, Legakis, Makarov, Tomić, S.B. Ćurčić, Mitic and Lucic, 2006, by original designation.

Distribution: EUROPE: Crete.

Ernstmayria apostolostrichasi Ćurčić and Dimitrijević

Ernstmayria apostolostrichasi Ćurčić and Dimitrijević, in Ćurčić, Dimitrijević, Legakis, Makarov, Tomić, S.B. Ćurčić, Mitić and Lučić, 2006: 39-42, figs 1-28; Ćurčić, Dimitrijević, Trichas, Tomić and Ćurčić, 2007: 758-764, figs 8-42.

 

Type locality: near Ierápetra, Lasíthi, Crete, Greece.

Distribution: Greece (Crete).

Ernstmayria venizelosi Ćurčić, Dimitrijević and Trichas

Ernstmayria venizelosi Ćurčić, Dimitrijević and Trichas, in Ćurčić, Dimitrijević, Trichas, Tomić and Ćurčić, 2007: 753-758, figs 1-7.

 

Type locality: Rouvas, Lasíthi, Crete, Greece.

Distribution: Greece (Crete).

Genus Microbisium Chamberlin

Microbisium Chamberlin, 1930: 20; Beier, 1932a: 136; Beier, 1939f: 186; Hoff, 1949b: 444-445; Hoff, 1956b: 2; Hoff, 1961: 431-432; Beier, 1963b: 202-203; Muchmore, 1979a: 194; Harvey, 1991a: 337;Muchmore, 1992: 131.

Afrobisium Beier, 1932a: 140 (synonymised by Mahnert, 1981a: 536).

Nepalobisium Beier, 1974b: 263-264; Harvey, 1991a: 391 (synonymised by Dashdamirov and Golovatch, 2006: 300).

 

Type species: of Microbisium: Obisium brunneum Hagen, 1868b, by original designation.

of Afrobisium: Obisium dogieli Redikorzev, 1924a, by original designation.

of Nepalobisium: Nepalobisium franzi Beier, 1974b, by original designation.

Distribution: ASIA: widespread.

EUROPE: widespread.

AMERICAS: widespread.

AFRICA: widespread.

Microbisium brevifemoratum (Ellingsen)

Obisium (Obisium) brevifemoratum Ellingsen, 1903: 13-17; Lessert, 1911: 32-33, fig. 25; Kästner, 1928: 11-12, fig. 45.

Obisium brevifemoratum Ellingsen: Schenkel, 1928: 64, figs 20a-b; Schenkel, 1929a: 322-323; Cirdei and Gutu, 1959: 2-3, fig. 2.

Microbisium brevifemoratum (Ellingsen): Beier, 1932a: 138, figs 168, 171; Tumšs, 1934: 13; Vachon, 1934i: 412; Roewer, 1936: fig. 74; Roewer, 1937: 249; Lohmander, 1939b: 288-291, fig. 3; Lohmander, 1945: 8, figs 1, 3-6; Franz and Beier, 1948: 476; Kaisila, 1949a: 74; Kaisila, 1949b: 78-79, fig. 3; Beier, 1952e: 4; Beier and Franz, 1954: 456; Meinertz, 1962: 51-54, figs 28a, 31, 32a-b; Beier, 1963b: 203, fig. 215; Lehtinen, 1964: 284-285; Meinertz, 1964: 392, map 2; Ressl, 1965: 289; Weygoldt, 1966d: 13, 15; Kölzel, 1967: 200; Rafalski, 1967: 13; Cloudsley-Thompson, 1968: 126 (as Microbisium femoratum [sic]); Verner, 1971: fig. 41; Palmgren, 1973: 7; Klausen, 1975: 64; Schawaller, 1985a: 3-4; Jędryczkowski, 1987b: 141, map 5; Schaefer, 1988: 100; Schawaller and Dashdamirov, 1988: 25, figs 39, 54; Schawaller, 1989: 10, figs 22-23; Harvey, 1991a: 337-338; Dashdamirov and Schawaller, 1992b: 39, fig. 2; Gärdenfors and Wilander, 1992: 28, fig. 3c; Jędryczkowski, 1995: 510; Platen, Blick, Bliss, Drogla, Malten, Martens, Sacher and Wunderlich, 1995: 54; Schawaller, 1995a: 124; Schawaller, 1995b: 1056; Drogla and Blick, 1996: 37; Henderickx, 1996: 1-3; Legg, 1998: 2; Legg, 1999: 1; Henderickx, 2000c: 67, fig. 2; Legg, 2003c: 3; Drogla and Lippold, 2004: 18; Mahnert, 2004: 464; Dashdamirov, 2005: 240, figs 59, 60; Dashdamirov and Golovatch, 2006: 299-300, figs 1-2.

Nepalobisium franzi Beier, 1974b: 265, fig. 2; Schawaller, 1987c: 204-205, figs 6-8; Harvey, 1991a: 391; Beron, 2002: 35 (synonymised by Dashdamirov and Golovatch, 2006: 300).

 

Type locality: of Microbisium brevifemoratum: Risør, Aust-Agder, Norway.

of Nepalobisium franzi: Taksang, above Tukche, Thakkhola, Nepal.

Distribution: Austria; Belgium; Azerbaijan; China (Hopeh, Ilan, Shaanxi); Denmark; Finland; Germany; Latvia; Nepal; Norway; Pakistan; Poland; Russia (Krasnoyarskiy Kray, Murmanskaya Oblast’); Sweden; Switzerland; United Kingdom.

Microbisium brevipalpe (Redikorzev)

Obisium brevipalpe Redikorzev, 1922a: 270-272, figs 12-14.

Microbisium brevipalpe (Redikorzev): Beier, 1932a: 138-139, fig. 172; Roewer, 1937: 249; Mani, 1962: 235 (as Microbisium brevipalpi [sic]); Harvey, 1991a: 338; Schawaller, 1995a: 124.

 

Type locality: Malyi Javinskij, Kamchatskaya, Russia.

Distribution: Russia (Kamchatskaya).

Microbisium brunneum (Hagen)

Obisium brunneum Hagen, 1868b: 52; Hagen, 1870: 269; Banks, 1890: 152 (as Obisium bruneum [sic]); Banks, 1895a: 12; Banks, 1908b: 4; Coolidge, 1908: 113; Ewing, 1911: 76, 79, fig. 13.

Obisium parvulum Banks, 1895a: 12 (misidentification, in part).

Microbisium brunneum (Hagen): Chamberlin, 1930: 20-21; Beier, 1932a: 139-140, fig. 174; Roewer, 1937: 250; Pearse, 1943: 464; Hoff, 1944b: 125-128, figs 1e-f, 2 (in part; see Microbisium parvulum (Banks)); Hoff, 1945a: 109; Hoff, 1945i: 323; Hoff, 1946g: 494-495; Pearse, 1946: 257; Proctor, 1946: 510; Rapp, 1946: 197; Hoff, 1949b: 445, figs 24a-b; Gering, 1956: 50; Hoff and Bolsterli, 1956: 161; Hoff, 1958: 8-9; Manley, 1969: 6-7, fig. 5; Muchmore, 1971a: 91; Nelson, 1975: 280, figs 18-19; Beier, 1976c: 1; Aitchison, 1979: 85; Nelson, 1984: 345; Sharkey, 1987: 17; Koponen and Sharkey, 1989: 388-389; Muchmore, 1990a: 513; Harvey, 1991a: 338; Snider and Nelson, 1991: 260; Blades and Marshall, 1994: 235; Finnamore, 1994: 220; Koponen, 1994: 55; Buddle, 2005: 14, 15, unnumbered fig.; Watermolen, 2007: 2.

Neobisium brunneum (Hagen): Brimley, 1938: 497.

Microbisium near brunneum (Hagen): McClure, 1943: 12.

Microbisium sp. prob. brunneum (Hagen): Carter and Brown, 1973: 1067.

Not Microbisium brunneum (Hagen): Hoff, 1945c: 34 (misidentification; see Microbisium parvulum (Banks)).

 

Type locality: Massachusetts, U.S.A.

Distribution: Canada (Alberta, Manitoba, Newfoundland, Nova Scotia, Quebec, Ontario); Costa Rica; U.S.A. (District of Columbia, Georgia, Illinois, Iowa, Maine, Massachusetts, Michigan, Missouri, New Jersey, New York, North Carolina, Ohio, South Carolina, Virginia, Utah, Wisconsin).

Microbisium congicum Beier

Microbisium congicum Beier, 1955a: 8-9, fig. 4; Beier, 1959d: 23; Harvey, 1991a: 338.

 

Type locality: Mukana, Parc National de l'Upemba, Shaba, Democratic Republic of Congo (Zaire).

Distribution: Democratic Republic of Congo (Zaire).

Microbisium dogieli (Redikorzev)

Obisium dogieli Redikorzev, 1924a: 192-193, figs 5-6.

Afrobisium dogieli (Redikorzev): Beier, 1932a: 140-141; Roewer, 1937: 250; Vachon, 1934i: 412-413.

Microbisium perpusillum Beier, 1955c: 532-533, fig. 4; Beier, 1959d: 23; Beier, 1962f: 14 (synonymised by Mahnert, 1981a: 536).

Microbisium dogieli (Redikorzev): Mahnert, 1981a: 536-537; Harvey, 1991a: 338; Beron, 2002: 35.

 

Type locality: of Obisium dogieli: Mabira, Kagera, Tanzania.

of Microbisium perpusillum: Mt Elgon, E. side, Rift Valley, Kenya.

Distribution: Democratic Republic of Congo (Zaire); Kenya; Rwanda; Tanzania.

Microbisium fagetum Cîrdei, Bulimar and Malcoci

Microbisium fagetum Cîrdei, Bulimar and Malcoci, 1967: 239-240, fig. 6; Harvey, 1991a: 339.

 

Type locality: Masivul Repedea, Iasi, Romania.

Distribution: Romania.

Microbisium lawrencei Beier

Microbisium lawrencei Beier, 1964k: 33-34, fig. 2; Harvey, 1991a: 339.

 

Type locality: Vumba, Zimbabwe.

Distribution: Zimbabwe.

Microbisium manicatum (L. Koch)

Obisium manicatum L. Koch, 1873: 61-62; L. Koch, 1878: 38; Pavesi, 1878: 363; Simon, 1885a: 349; Ellingsen, 1909a: 214; Krausse-Heldrungen, 1912: 65; Daday, 1918: 2; Navás, 1925: 124.

Obisium (Obisium) manicatum L. Koch: Daday, 1888: 132, 188, plate 4 fig. 25; Ellingsen, 1910a: 398-399; Ellingsen, 1912c: 173; Nonidez, 1917: 40; Jeannel, 1926: 152; Beier, 1928: 296; Beier, 1929d: 156.

Microbisium manicatum (Ellingsen): Beier, 1932a: 137, fig. 169; Roewer, 1937: 249; Wolf, 1938: 619; Beier, 1939f: 186; Caporiacco, 1951e: 69; Beier, 1963b: 205, fig. 217; Kobakhidze, 1965b: 541; Kobakhidze, 1966: 704; Lazzeroni, 1969a: 333; Lazzeroni, 1969b: 236; Cîrdei, Bulimar and Malcoci, 1970: 11-12, fig. 5; Lazzeroni, 1970a: 207; Harvey, 1991a: 339; Dashdamirov and Schawaller, 1992b: 39; Gardini, 2000: 96; Delfosse, 2003: 37; Zaragoza, 2007a: 25.

 

Type locality: Greece.

Distribution: France (mainland); Georgia; Greece (mainland, Kyklades); Hungary; Italy (mainland, Sardinia); Romania; Spain (mainland).

Microbisium parvulum (Banks)

Obisium parvulum Banks, 1895a: 12 (in part; see Microbisium brunneum (Hagen)); Banks, 1904a: 141; Coolidge, 1908: 113; Ellingsen, 1909b: 220.

Microbisium parvulum (Banks): Chamberlin, 1930: 21-22; Chamberlin, 1931a: figs 28h, 35n; Beier, 1932a: 139, fig. 173; Vachon, 1934i: 412; Roewer, 1937: 250; Brimley, 1938: 497; Hoff, 1946g: 495-496; Hoff, 1956b: 3-4; Hoff and Bolsterli, 1956: 161; Hoff, 1958: 9; Hoff, 1959b: 4, etc.; Hoff, 1961: 432, figs 7-9; Rowland and Reddell, 1976: 15; Nelson, 1984: 343-349, maps 1-2; Corey and Taylor, 1987: 165; Muchmore, 1990a: 513; Harvey, 1991a: 339; Shultz, 1991: fig. 6c; Snider and Nelson, 1991: 260; Muchmore, 1992: 131; Buddle, 2005: 15; Brand, 2006: 127; Francke and Villegas-Guzmán, 2006: 295; Watermolen, 2007: 3.

Microbisium brunneum (Hagen): Hoff, 1944b: 125-128, figs 1e-f, 2 (in part) (misidentification); Hoff, 1945c: 34 (misidentification).

Microbisium confusum Hoff, 1946g: 496-497; Hoff, 1949b: 446, figs 25a-c; Hoff, 1956b: 3; Hoff and Bolsterli, 1956: 160; Hoff, 1958: 9; Hoff, 1961: 432-433, figs 10-11; Hoff, 1963a: 2-3; Wheeler and Wheeler, 1964: 8; Lawson, 1968: 4351; Lawson, 1969: 3-4, figs 1-5; Manley, 1969: 6, fig. 4; Weygoldt, 1969a: fig. 105; Muchmore, 1971a: 91; Knowlton, 1972: 2; Muchmore, 1972d: 110; Nelson, 1973: 232-235, figs a-c; Knowlton, 1974: 3; Nelson, 1975: 280-281, figs 20-24; W.F. Rapp, 1978a: 5-7; W.F. Rapp, 1978b: 12; Aitchison, 1979: 85; Muchmore, 1979a: 194; Muchmore, 1981f: 381; Nelson, 1982: 262-274, figs 1-28; W.F. Rapp, 1986: 220-221; Sharkey, 1987: 17; D.W. Zeh, 1987b: 1086 (synonymised by Nelson, 1984: 349).

Microbisium sp. prob. confusum Hoff: Graves and Graves, 1969: 267.

 

Type locality: of Obisium parvulum: North America [Hoff (1958) invalidly declared Bernalillo County, New Mexico, as type locality].

of Microbisium confusum: Antioch, Lake County, Illinois, U.S.A.

Distribution: Canada (Manitoba, Nova Scotia, Ontario, Quebec); Costa Rica; El Salvador; Mexico; U.S.A. (Arkansas, California, Colorado, Connecticut, Florida, Georgia, Idaho, Illinois, Indiana, Iowa, Kansas, Kentucky, Maine, Maryland, Massachusetts, Michigan, Minnesota, Mississippi, Missouri, Nebraska, New Hampshire, New Jersey, New Mexico, New York, North Carolina, North Dakota, Ohio, Oklahoma, Pennsylvania, South Carolina, South Dakota, Tennessee, Texas, Utah, Vermont, Virginia, Wisconsin).

Microbisium pygmaeum (Ellingsen)

Obisium pygmaeum Ellingsen, 1907c: 12-14.

Microbisium pygmaeum (Ellingsen): Beier, 1932a: 140; Roewer, 1937: 250; Morikawa, 1952b: 254-255, fig. 9; Kobari, 1984a: 57-62, figs 1, 2a-f, 3a-e; Sato, 1985: 22, figs 1-2; Sato, 1988: 12; Sato, Takano and Nakajima, 1988: 23; Sakayori, 1989: 56-61, figs 1-27; Sakayori, 1990: 33, fig. 2B; Harvey, 1991a: 340; Nakajima, Takano, Sato and Tashiro, 1991: 16; Sato, 1991a: 23, fig. 68-2; Hong and T.H. Kim, 1993: 2; Sato, 1994b: 80; Sakayori, 1995: 28, fig. D; Nunomura, Hirauchi and Sato, 1997: 12-13; Sakayori, 1998b: 706, fig. 3; Nunomura, Suma, Ishii and Sato, 1997: 86; Sakayori, 1998a: 300; Sato and Yamamoto, 2000: 666; Sato and Yamauchi, 2001: 63-65.

Neobisium (Parobisium) pygmaeum (Ellingsen): Morikawa, 1960: 114-115, plate 2 fig. 3; Sato, 1979a: 94; Sato, 1979d: 43; Sato, 1982c: 32; Kobari, 1983: 65-70, fig. 1.

Neobisium pygmaeum (Ellingsen): Sato, 1979b: 13, figs 1-3, 5; Sato, 1983b: 16, figs 2, 4; Sato, 1984c: 14, figs 1, 3; Sato, 1984d: fig. 2; W.K. Lee, 1981: 130, figs 2a-b.

 

Type locality: Negishi, S. of Yokohama, Kanagawa, Japan; Okayama, Okayama Prefecture, Japan; and Kanagawa, N. of Yokohama, Kanagawa Prefecture, Japan.

Distribution: Japan; South Korea.

Microbisium suecicum Lohmander

Microbisium parvulum suecicum Lohmander, 1945: 7-8, figs 2, 7-10.

Microbisium suecicum Lohmander: Meinertz, 1962: 54-55, figs 4c, 28b-c, 33a; Beier, 1963b: 204; Rafalski, 1967: 13; Gardini, 1976a: 251-255, figs 1-8; Mahnert, 1983d: 362; Gardini, 1985b: 60; Jędryczkowski, 1987b: 142, map 6; Mahnert, 1990b: 683; Harvey, 1991a: 340; Gärdenfors and Wilander, 1992: 28-29; Drogla and Lippold, 1994: 75; Gardini, 1995a: 5; Platen, Blick, Bliss, Drogla, Malten, Martens, Sacher and Wunderlich, 1995: 54; Drogla and Blick, 1996: 37; Henderickx, 1999a: 65-67, figs 1-3; Gardini, 2000: 96; Henderickx, 2000c: 67; Šťáhlavský and Ducháč, 2001: 47; Drogla and Lippold, 2004: 18, fig. 2.4; Mahnert, 2004: 464; Šťáhlavský and Ducháč, 2005: 83; Šťáhlavský, 2006a: 170.

Microbisium cfr. suecicum Lohmander: Callaini, 1988b: 42-46, figs 3c-m.

 

Type locality: several localities in Sweden.

Distribution: Austria; Belgium; Czech Republic; Germany; Hungary; Italy (mainland); Morocco; Poland; Sweden; Switzerland.

Microbisium zariquieyi (Navás)

Obisium zariquieyi Navás, 1919: 212-214, figs 9a-c; Navás, 1925: 124, figs 17a-c.

Microbisium (?) zariquieyi (Navás): Beier, 1932a: 140; Roewer, 1937: 250.

Microbisium zariquieyi (Navás): Beier, 1939f: 187-188, fig. 17; Beier, 1959f: 122; Beier, 1963b: 204-205, fig. 216; Harvey, 1991a: 340; García Carrillo, 1998: 12; Zaragoza, 2007a: 25.

 

Type locality: Santa Fé del Montseny, Barcelona, Cataluña, Spain.

Distribution: Spain (mainland).

Genus Neobisium Chamberlin

Obisium Illiger: Walckenaer, 1802: 252-253; Risso, 1826: 158; Théis, 1832: 63; C.L. Koch, 1839: 5; C.L. Koch, 1850: 5; Menge, 1855: 26; McIntire, 1868: 9; Stecker, 1874b: 239; Stecker, 1875d: 89; Simon, 1879a: 51-52; Tömösváry, 1882b: 219-220; Canestrini, 1885: no. 4; Daday, 1888: 127; O.P.-Cambridge, 1892: 212; Becker, 1896: 332-333; Tullgren, 1899a: 180; Planet, 1905: 285-286; Tullgren, 1906a: 204; With, 1906: 76; Kew, 1911a: 52; Lessert, 1911: 27; Pratt, 1927: 410.

Obisium (Obisium) Illiger: Kew, 1911a: 54; Lessert, 1911: 29.

Neobisium Chamberlin, 1930: 11-12; Beier, 1932a: 78; Beier, 1939d: 26; Chamberlin, 1935b: 480; Roewer, 1937: 245; Beier, 1939f: 167; Hoff, 1949b: 444; Petrunkevitch, 1955: 81; Morikawa, 1960: 112; Hoff, 1961: 427; Beier, 1963b: 81; Ćurčić, 1978b: 132; Legg and Jones, 1988: 75; Harvey, 1991a: 346.

 

Type species: Obisium muscorum Leach, 1817 (junior synonym of Chelifer carcinoides Hermann, 1804), by original designation.

Distribution: EUROPE: widespread

NORTH AMERICA: introduced

Subgenus Neobisium (Blothrus) Schiödte

Blothrus Schiödte, 1847: 80; Schiödte, 1851a: 23; Schiödte, 1851b: 148-149; Simon, 1872: 223-224; Tömösváry, 1882b: 233; Planet, 1905: 286; Chamberlin, 1930: 9; Harvey, 1985a: 367.

Obisium (Blothrus) Schiödte: Daday, 1888: 127; Nonidez, 1925: 45-53.

Neobisium (Blothrus) Schiödte: Beier, 1932a: 106; Beier, 1939d: 36-37; Beier, 1939f: 174-175; Harvey, 1991a: 373.

Blothrus (Blothrus) Schiödte: Harvey, 1985a: 367.

 

Type species: Blothrus spelaeus Schiödte, 1847, by monotypy.

Neobisium (Blothrus) abeillei (Simon)

Blothrus abeillei Simon, 1872: 224-226, fig. 10; L. Koch, 1873: 43-44; Planet, 1905: 287, unnumbered fig. on p. 287; Berland, 1932: fig. 84.

Obisium (Blothrus) abeillei (Simon): Simon, 1879a: 68-69, 312, plate 19 fig. 12; Ellingsen, 1912c: 167; Nonidez, 1917: 32; Jeannel, 1926: 154-155; Berland, 1929: 17.

Obisium abeillei (Simon): Ellingsen, 1908a: 416; Navás, 1925: 120.

Blothrus abeilli Simon: Chamberlin, 1930: 11; Chamberlin, 1931a: figs 28i, 35a-b; Vachon, 1936a: 78.

Neobisium (Blothrus) abeillei (Simon): Beier, 1932a: 107-108, fig. 124; Roewer, 1937: 248; Vachon, 1947c: figs 1, 4, 7, 9; Chamberlin and Malcolm, 1960: fig. 1d; Beier, 1963b: 138-139, fig. 139; Heurtault, 1986c: 26, fig. 1; Harvey, 1991a: 373; Delfosse, 2003: 34; Zaragoza, 2007a: 66.

Neobisium abeilli (Simon): Wolf, 1938: 609.

Not Obisium abeillei (Simon): Lebedinsky, 1904: 80, fig. 24 (misidentification; true identification unknown); Gozo, 1908: 129 (not seen) (misidentification; see Roncus gestroi Beier).

Not Obisium abeillei (Simon): Gozo, 1908: 129-130 (misidentification; see Roncus gestroi Beier).

 

Type locality: Grotte d'Estellas, Ariège, Midi-Pyrénées, France.

Distribution: France (mainland).

Neobisium (Blothrus) absoloni Beier

Neobisium (Blothrus) absoloni Beier, 1938a: 6; Harvey, 1991a: 373.

 

Type locality: Vilina Jama, Lebršnik Planina, Gatačkog Polje, Republika Srpska, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina; Yugoslavia (Montenegro).

Neobisium (Blothrus) absoloni absoloni Beier

Neobisium (Blothrus) absoloni absoloni Beier: Beier, 1939d: 48-49, fig. 52; Beier, 1963b: 171, fig. 179; Ćurčić, 1974a: 18.

 

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) absoloni grande Beier

Neobisium (Blothrus) absoloni grande Beier, 1939d: 49-50, fig. 55; Beier, 1963b: 172; Ćurčić, 1974a: 18-19; Harvey, 1991a: 373; Dimitrijević, 2000c: 39; Ćurčić, Dimitrijević and Legakis, 2004: 240.

 

Type locality: Sertovom Dolu (as Jama in Sertov), E. of Gacko Polje, Montenegro, Yugoslavia.

Distribution: Bosnia-Herzegovina; Yugoslavia (Montenegro).

Neobisium (Blothrus) absoloni tacitum Beier

Neobisium (Blothrus) absoloni tacitum Beier, 1939d: 49, fig. 54; Beier, 1963b: 172; Ćurčić, 1974a: 19; Harvey, 1991a: 374.

 

Type locality: Borija Pećina, near Krbljina, Treskavici Planini, Republika Srpska, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) alae Ćurčić, Dimitrijević, S.B. Ćurčić and Mitić

Neobisium alae Ćurčić, Dimitrijević, S.B. Ćurčić and Mitić, 2004: 84-88, figs 1-8; Ćurčić, Dimitrijević and Legakis, 2004: 241.

 

Type locality: Lepetane, near Kotor, Montenegro, Yugoslavia.

Distribution: Yugoslavia (Montenegro).

Neobisium (Blothrus) albanicum (G. Müller)

Obisium (Blothrus) albanicum G. Müller, 1931: 126.

Obisium albanicum G. Müller: Wolf, 1938: 619.

Neobisium (Blothrus) albanicum (G. Müller): Beier, 1939d: 68-69, fig. 82; Beier, 1963b: 163, fig. 170; Beier, 1970f: 320; Harvey, 1991a: 374.

 

Type locality: Berat Cave, Paftali, Berat, Albania.

Distribution: Albania.

Neobisium (Blothrus) albanorum Ćurčić, Dimitrijević, Rađa and Vujčić-Karlo

Neobisium albanorum Ćurčić, Dimitrijević, Rađa and Vujčić-Karlo, 2006: 3-10, figs 1-30.

 

Type locality: Pusi I Zylihasë Cave, near Gjirokaster, Albania.

Distribution: Albania.

Neobisium (Blothrus) auberti Leclerc

Neobisium (Blothrus) auberti Leclerc, 1982a: 40-43, figs 1-7, plates 1-2; Heurtault, 1986c: 26; Harvey, 1991a: 374; Delfosse, 2003: 35.

 

Type locality: Sadoux à Pradelle, Drôme, Rhône-Alpes, France.

Distribution: France (mainland).

Neobisium (Blothrus) aueri Beier

Neobisium (Blothrus) aueri Beier, 1962i: 2-3, fig. 1; Beier, 1963b: 172-173, fig. 180; Beier, 1970f: 320; Thaler, 1980: 391; Gaisberger, 1984b: 57; Christian, 1986: 122, fig. 1; Harvey, 1991a: 374; Ducháč, 1999b: 175-180, fig. 5; Ducháč, 2004c: 54-55, fig. 8; Mahnert, 2004: 464.

 

Type locality: Almberg-Eis and Tropfstein Cave, near Grundl-see, Steiermark, Austria.

Distribution: Austria.

Neobisium (Blothrus) babusnicae Ćurčić

Neobisium babusnicae Ćurčić, 1980c: 249-254, figs 1-6; Ćurčić, 1983b: fig. 2; Ćurčić, 1988e: fig. 1.

Neobisium (Blothrus) babusnicae Ćurčić: Harvey, 1991a: 374; Ćurčić, 1996: 344; Dimitrijević, 2000c: 39; Ćurčić, Dimitrijević and Legakis, 2004: 241

 

Type locality: Pećina Pripor Cave, Resnik, near Babušnica, Serbia, Yugoslavia.

Distribution: Yugoslavia (Serbia).

Neobisium (Blothrus) birsteini (Lapschoff)

Blothrus birsteini Lapschoff, 1940: 62-63, 73, figs 1-2; Kobakhidze, 1965b: 541; Kobakhidze, 1966: 703.

Neobisium (? Blothrus) birsteini (Lapschoff): Mahnert, 1979b: 263-264, figs 7-12.

Neobisium (Blothrus) birsteini (Lapschoff): Harvey, 1991a: 374; Dashdamirov and Schawaller, 1992b: 40, 49.

 

Type locality: Cave Tarkiladze (as Höhle Tarkyladze), near Gudauten, Sokhumi (as Sukhumi), Abkhazia, Georgia.

Distribution: Georgia.

Neobisium (Blothrus) bolivari (Nonidez)

Obisium (Blothrus) bolivari Nonidez, 1917: 32-36, figs 8, 9a-c; Nonidez, 1925: 72-73, figs 11a-b.

Obisium bolivari Nonidez: Navás, 1925: 120.

Neobisium bolivari (Nonidez): Wolf, 1938: 609.

Neobisium (Blothrus) bolivari (Nonidez): Beier, 1932a: 116, fig. 140; Roewer, 1937: 248; Beier, 1939f: 175; Beier, 1963b: 144-145, fig. 146; Beier, 1970f: 320; Mahnert, 1977c: 81-83, figs 32-33; Estany, 1980b: 526, map 1; Zaragoza, 1986: fig. 1; Harvey, 1991a: 374; Zaragoza, 2007a: 30.

 

Type locality: Cueva de Albia, Sierra Salvada, Burgos, Castilla y León, Spain.

Distribution: Spain (mainland).

Neobisium (Blothrus) boneti Beier

Neobisium (Blothrus) boneti Beier, 1931a: 11-12, fig. 3; Beier, 1932a: 107, fig. 123; Roewer, 1937: 247; Beier, 1939f: 175; Beier, 1963b: 137, fig. 137; Estany, 1980b: 526, map 1; Zaragoza, 1986: fig. 1; Bellés, 1987: fig. 39; Harvey, 1991a: 374-375; Zaragoza, 2007a: 30-31.

Neobisium boneti (Nonidez): Wolf, 1938: 609.

 

Type locality: Cueva de Mauloetxea (as Mauloechea), Abaurrea Alta, Navarra, Spain.

Distribution: Spain (mainland).

Neobisium (Blothrus) breuili (Bolivar)

Obisium breuili Bolivar, 1924: 103-104, fig. 2; Navás, 1925: 121.

Obisium (Blothrus) breuili Bolivar: Nonidez, 1925: 79-81, figs 17, 18a-b.

Neobisium (Blothrus) breuili (Bolivar): Beier, 1932a: 119-120, fig. 146; Roewer, 1937: 248; Beier, 1939f: 177; Beier, 1963b: 140-141, fig. 142; Beier, 1970f: 320; Estany, 1980b: 527, map 2 (as Neobisium (Blothrus) breuli [sic]); Zaragoza, 1986: fig. 1; Bellés, 1987: figs 40, 42; Harvey, 1991a: 375; Zaragoza, 2007a: 31.

Neobisium breuili (Bolivar): Wolf, 1938: 609.

 

Type locality: Cueva de Martinchurito I, Lecumberri, Navarra, Spain.

Distribution: Spain (mainland).

Neobisium (Blothrus) brevimanum (J. Frivaldsky)

Blothrus brevimanum J. Frivaldsky, 1865b: 80, fig. 3 (not seen).

Obisium (Obisium) brevipes (J. Frivaldsky): Beier, 1928: 306 (in part).

Neobisium (Blothrus) (?) brevimanum (J. Frivaldsky): Beier, 1932a: 124.

Neobisium (Blothrus) brevimanum (J. Frivaldsky): Harvey, 1991a: 375.

 

Type locality: "Karpathen".

Distribution: "Karpathen".

Neobisium (Blothrus) brevipes (J. Frivaldsky)

Blothrus brevipes J. Frivaldsky, 1865a: 38-40; Heyden, 1869: 59; Daday, 1880a: 192; Tömösváry, 1882b: 234-235; Daday, 1918: 2.

Obisium (Blothrus) brevipes (J. Frivaldsky): Daday, 1888: 128, 184; Beier, 1928: 306 (in part; see Neobisium (Blothrus) brevimanum J. Frivaldsky and Neobisium (Blothrus) minutus Tömösváry); Beier, 1929a: 14a; Beier, 1929e: fig. 5b.

Neobisium (Blothrus) brevipes (J. Frivaldsky): Beier, 1932a: 110-111, fig. 130; Roewer, 1937: 248; Beier, 1939c: 14-15, fig. 9; Decou and Negrea, 1969: 522; Beier, 1970f: 320; Harvey, 1991a: 375; Ducháč, 1999b: 175-180, fig. 4; Ducháč, 2000: 7-10, figs 1-2.

Neobisium brevipes (J. Frivaldsky): Wolf, 1938: 610.

 

Type locality: Pesterea dela Ferice (as Fericsei barhang), near Ferice, Beius, Bihor, Romania; Pesterea Zemeilor din Onceasa (as Oncsásza barhang), near Budureasa, Beius, Bihor, Romania; and Pesterea dela Fanate (as

Fonáczai barhang), Fanate, Orodea, Romania.

Distribution: Romania.

Neobisium (Blothrus) brevipes brevipes (J. Frivaldsky)

Neobisium (Blothrus) brevipes brevipes (J. Frivaldsky): Beier, 1963b: 153-154.

 

Distribution: Romania.

Neobisium (Blothrus) brevipes montanum Beier

Neobisium (Neobisium) brevipes montanum Beier, 1939c: 15-16, fig. 10.

Neobisium (Blothrus) brevipes montanum Beier: Beier, 1963b: 154, fig. 159; Harvey, 1991a: 375.

 

Type locality: Pesterea dela Alun, Beius, Bihor, Romania.

Distribution: Romania.

Neobisium (Blothrus) caecum Beier

Neobisium (Neobisium) caecum Beier, 1939d: 35-36, fig. 41; Beier, 1947a: 173-174, fig. 1m.

Neobisium (Blothrus) caecum Beier: Beier, 1963b: 151, fig. 154; Ćurčić, 1974a: 19; Harvey, 1991a: 375; Ozimec, 2004: 387.

Neobisium caecum Beier: Ozimec, in Gottstein Matočec et al., 2002: 55.

 

Type locality: Planini Velika Žaba, SW. of Popovo Polje, Federation of Bosnia and Herzegovina, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina; Croatia.

Neobisium (Blothrus) carnae Beier

Neobisium (Blothrus) carnae Beier, 1938a: 7; Beier, 1970f: 320; Harvey, 1991a: 375.

 

Type locality: Koaca, Radoš Planina, N. of Popovo Polje, Federation of Bosnia and Herzegovina, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) carnae carnae Beier

Neobisium (Blothrus) carnae carnae Beier: Beier, 1939d: 61-62, fig. 73; Beier, 1963b: 148, fig. 151; Ćurčić, 1974a: 18.

 

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) carnae fraternum Beier

Neobisium (Blothrus) carnae fraternum Beier, 1939d: 62, fig. 74; Beier, 1963b: 148-149; Ćurčić, 1974a: 19; Harvey, 1991a: 376.

 

Type locality: Dvogrlama, Radoš Planina, Federation of Bosnia and Herzegovina, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) casalei Gardini

Neobisium (Blothrus) casalei Gardini, 1985a: 57-60, figs 10-16; Harvey, 1991a: 376.

 

Type locality: Drako Trypa Cave, Vytina (as Vitina), Arkadia, Greece.

Distribution: Greece (mainland).

Neobisium (Blothrus) cervelloi Mahnert

Neobisium (Blothrus) cervelloi Mahnert, 1977c: 78-80, figs 25-31; Zaragoza, 1986: fig. 1; Harvey, 1991a: 376; García Carrillo, 1998: 12; Zaragoza, 2007a: 31.

Neobisium (Blothrus) aff. cervalloi Mahnert: Mahnert, 1977c: 80-81; Zaragoza, 1986: fig. 1.

 

Type locality: Avenc T-1, Sierra Arañonera, Monte Perdido, Torla, Huesca, Aragón, Spain.

Distribution: Spain (mainland).

Neobisium (Blothrus) chaimweizmanni Ćurčić and Dimitrijević

Neobisium chaimweizmanni Ćurčić and Dimitrijević, in Ćurčić, Dimitrijević, S.B. Ćurčić, Tomić and N.B. Ćurčić, 2004: 85-88, figs 1-7; Ćurčić, Dimitrijević and Legakis, 2004: 244; Ozimec, 2004: 387; Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 77.

 

Type locality: Trojama Jama (as Trojama Pit), Split, Mosor Planina, Splitsko-Dalmatinska, Croatia.

Distribution: Croatia.

Neobisium (Blothrus) closanicum Dumitresco and Orghidan

Neobisium (Blothrus) closanicus Dumitresco and Orghidan, 1970b: 104-111, figs 15-25 (as Neobisium (Blothrus) cloşanicus); Harvey, 1991a: 376.

 

Type locality: Peştera Cloşani, Olt, Romania.

Distribution: Romania.

Neobisium (Blothrus) coiffaiti Heurtault

Neobisium (Blothrus) coiffaiti Heurtault, 1986c: 26, fig. 4; Harvey, 1991a: 376; Delfosse, 2003: 35.

 

Type locality: Grotte de la Palle, Rieulhes, Hautes-Pyrénées, Midi-Pyrénées, France.

Distribution: France (mainland).

Neobisium (Blothrus) creticum (Beier)

Obisium (Blothrus) creticum Beier, 1931d: 92-93, figs 1, 3a; Wolf, 1938: 610.

Neobisium (Blothrus) creticum (Beier): Beier, 1932a: 108-109, fig. 126; Roewer, 1937: 248; Beier, 1939d: 45; Beier, 1963b: 161-162, fig. 168; Harvey, 1991a: 376.

 

Type locality: Katholiko Cave, near Akrotiri, Crete, Greece.

Distribution: Greece (mainland, Crete).

Neobisium (Blothrus) dalmatinum Beier

Neobisium (Blothrus) dalmatinum Beier, 1938a: 7; Harvey, 1991a: 376; Ozimec, 2004: 387.

Neobisium (Blothrus) dalmatinum dalmatinum Beier: Beier, 1939d: 51-52, figs 57-58; Beier, 1963b: 149, fig. 152; Beier, 1970f: 320; Ćurčić, 1974a: 19.

Neobisium (Blothrus) dalmatinum aberrans Beier, 1938a: 7; Beier, 1939d: 52, fig. 59; Beier, 1963b: 149-150; Ćurčić, 1974a: 19 (synonymised by Ćurčić, 1988b: 43).

Neobisium dalmatinum Beier: Ćurčić, 1988b: 43-47, figs 86-125; Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 13.

Neobisium dalmatinum aberrans Beier: Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 56.

 

Type locality: of Neobisium (Blothrus) dalmatinum dalmatinum: Balićeva Špilja (as Kraljeva Pećina), Split (as Spalato), Splisko-Dalmatinska, Croatia.

of Neobisium (Blothrus) dalmatinum aberrans: Sniježnica pod Ljubljanom (as Kraljeva Pečina), Ljubljan, Spilt, Mosor Planina, Splitsko-Dalmatinska, Croatia.

Distribution: Croatia.

Neobisium (Blothrus) davidbengurioni Ćurčić and Dimitrijević

Neobisium davidbengurioni Ćurčić and Dimitrijević, in Ćurčić, Dimitrijević, S.B. Ćurčić, Tomić and N.B. Ćurčić, 2004: 88-93, figs 8-23; Ćurčić, Dimitrijević and Legakis, 2004: 244.

 

Type locality: Jama u Vjetrenim Pit, Vjetrena Brda, Mt Durmitor, Montenegro, Yugoslavia.

Distribution: Yugoslavia (Montenegro).

Neobisium (Blothrus) deschmanni (G. Joseph)

Obisium deschmanni G. Joseph, 1882: 22.

Neobisium (Blothrus) (?) deschmanni (G. Joseph): Beier, 1932a: 124; Roewer, 1937: 248.

Neobisium deschmanni (G. Joseph): Wolf, 1938: 611.

Neobisium (Blothrus) deschmanni (G. Joseph): Harvey, 1991a: 377.

 

Type locality: Grotte von Luëg, Tirol, Austria

Distribution: Austria.

Neobisium (Blothrus) dinaricum Hadži

Neobisium (Blothrus) dinaricum Hadži, 1933b: 158-165, figs 18a-c, 19a-c, 20a-d, 21a-b; Hadži, 1933c: 186-189; Roewer, 1937: 248; Beier, 1970f: 320; Harvey, 1991a: 377; Ćurčić, Dimitrijević and Legakis, 2004: 244-245; Ozimec, 2004: 387.

 

Type locality: Crkvice, Krivošije, Montenegro, Yugoslavia.

Distribution: Croatia; Yugoslavia (Montenegro).

Neobisium (Blothrus) dinaricum caligatum Beier

Neobisium (Blothrus) dinaricum caligatum Beier, 1938a: 7; Beier, 1939d: 54-55, fig. 63; Beier, 1963b: 159, fig. 164; Ćurčić, 1974a: 19; Harvey, 1991a: 377; Dimitrijević, 2000c: 39; Ćurčić, Dimitrijević and Legakis, 2004: 245.

 

Type locality: Doluja Špilja, Dvršnik, Krivošije, Montenegro, Yugoslavia.

Distribution: Yugoslavia (Montenegro).

Neobisium (Blothrus) dinaricum dinaricum Hadži

Neobisium (Blothrus) dinaricum dinaricum Hadži: Beier, 1939d: 53; Beier, 1963b: 157-158; Ćurčić, 1974a: 19; Dimitrijević, 2000c: 39.

 

Distribution: Yugoslavia (Montenegro).

Neobisium (Blothrus) dinaricum tartareum Beier

Neobisium (Blothrus) dinaricum tartareum Beier, 1938a: 7; Beier, 1939d: 53-54, fig. 62; Beier, 1963b: 158-159; Ćurčić, 1974a: 20; Harvey, 1991a: 377; Dimitrijević, 2000c: 39; Ćurčić, Dimitrijević and Legakis, 2004: 245.

 

Type locality: Napoda, Krivošije, Montenegro, Yugoslavia.

Distribution: Yugoslavia (Montenegro).

Neobisium (Blothrus) georgecastriotae Ćurčić, Dimitrijević, Rađa, Dudić, Šimić and Vujčić-Karlo

Neobisium stitkovense Ćurčić, Dimitrijević, Rađa, Dudić, Šimić and Vujčić-Karlo, 2006: 101-103, figs 1-8.

 

Type locality: Pusi i Kerpajt Pit, Progomat, Albania.

Distribution: Albania.

Neobisium (Blothrus) goldameirae Ćurčić and Dimitrijević

Neobisium goldameirae Ćurčić and Dimitrijević, in Ćurčić, Dimitrijević, S.B. Ćurčić, Tomić and N.B. Ćurčić, 2004: 97-103, figs 39-60; Ćurčić, Dimitrijević and Legakis, 2004: 247.

 

Type locality: Arapova Pećina Cave, Grabovica, Mt Durmitor, Montenegro, Yugoslavia.

Distribution: Yugoslavia (Montenegro).

Neobisium (Blothrus) golemanskyi Ćurčić and Dimitrijević

Neobisium golemanskyi Ćurčić and Dimitrijević, 2001: 148-154, figs 7-18; Ćurčić, Dimitrijević and Legakis, 2004: 247.

 

Type locality: Peštera Bozguni Cave, Patiška Reka, near Skopje, Skopje, Macedonia.

Distribution: Macedonia.

Neobisium (Blothrus) hadzii Beier

Neobisium (Blothrus) hadzii Beier, 1938a: 6; Beier, 1939d: 45, fig. 49; Beier, 1963b: 174-175, fig. 183; Ćurčić, 1974a: 20; Harvey, 1991a: 377; Ozimec, 2004: 387 (as Neobisium (Neobisium) hadžii).

Neobisium hadzii Beier: Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 33.

 

Type locality: Jezero Špilja (as Jezero Pečina), Popovo Polje, Duprovačko-Neretvanska, Croatia.

Distribution: Croatia.

Neobisium (Blothrus) heros Beier

Neobisium (Blothrus) heros Beier, 1938a: 6; Beier, 1939d: 40-41, fig. 42; Beier, 1963b: 147, fig. 149; Beier, 1970f: 320; Ćurčić, 1974a: 20; Harvey, 1991a: 377; Ćurčić, 1996: 344; Dimitrijević, 2000c: 39; Ozimec, 2004: 387.

Neobisium heros Beier: Ćurčić, Dimitrijević and Legakis, 2004: 247.

 

Type locality: Pećina u Kučericama, Orjenu, Montenegro, Yugoslavia.

Distribution: Croatia; Yugoslavia (Montenegro).

Neobisium (Blothrus) hians Mahnert

Neobisium (Blothrus) hians Mahnert, 1979b: 261-263, figs 1-6; Harvey, 1991a: 377.

 

Type locality: Grotte In Dagi, Dösemealth (as Dosemealti), Antalya, Turkey.

Distribution: Turkey.

Neobisium (Blothrus) hiberum Beier

Neobisium (Blothrus) hiberum Beier, 1931a: 13-14, fig. 5; Beier, 1932a: 115, fig. 137; Roewer, 1937: 248; Beier, 1939f: 175; Beier, 1963b: 136, fig. 135; Zaragoza, 1986: fig. 1; Harvey, 1991a: 377; Zaragoza, 2007a: 31.

Neobisium hiberum Beier: Wolf, 1938: 611.

 

Type locality: Sima de la Raya, Tamajón, Guadalajara, Castilla-La Mancha, Spain.

Distribution: Spain (mainland).

Neobisium (Blothrus) hypochthon Beier

Neobisium (Blothrus) hypochthon Beier, 1938a: 6; Beier, 1939d: 50-51, fig. 56; Beier, 1963b: 173, fig. 181; Vachon and Gabbutt, 1964: fig. 15; Ćurčić, 1974a: 20; Harvey, 1991a: 378; Dimitrijević, 2000c: 39.

 

Type locality: Pećina Babič, Popovo Polje, Federation of Bosnia and Herzegovina, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina; Yugoslavia (Montenegro).

Neobisium (Blothrus) imbecillum Beier

Neobisium (Blothrus) imbecillum Beier, 1938a: 6; Beier, 1939d: 44, fig. 47; Beier, 1963b: 169, fig. 176; Ćurčić, 1974a: 20; Harvey, 1991a: 378.

 

Type locality: Pećina Krbljina, Treskavica Planina, Republika Srpska, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) infernum Beier

Neobisium (Blothrus) infernum Beier, 1938a: 7; Beier, 1939d: 59-60, fig. 69; Beier, 1963b: 155, fig. 161; Ćurčić, 1974a: 20; Harvey, 1991a: 378.

 

Type locality: Jama kod Komasovice, near Krstač, Popovo Polje, Orjena, Federation of Bosnia and Herzegovina, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) insulare Beier

Neobisium (Blothrus) insulare Beier, 1938a: 7; Beier, 1939e: 311, fig. 6; Beier, 1963b: 161; Ćurčić, 1974a: 20; Mahnert, 1980g: 97; Christian and Potocnik, 1985: 17; Harvey, 1991a: 378; Ozimec, 2004: 387.

Neobisium insulare Beier: Ćurčić, 1988b: 49-50, figs 133-137; Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 15.

 

Type locality: Biserujka Špilja, Insel Deglia, Krk, Primorsko-Goranska, Croatia.

Distribution: Croatia.

Neobisium (Blothrus) jeanneli (Ellingsen)

Obisium (Blothrus) jeanneli Ellingsen, 1912c: 167-169; Nonidez, 1917: 36-37, figs 10a-c; Nonidez, 1925: 82-83, figs 19a-b.

Obisium jeanneli Ellingsen: Navás, 1925: 121.

Neobisium (Blothrus) jeanneli (Ellingsen): Beier, 1932a: 120, fig. 147; Roewer, 1937: 248; Beier, 1939f: 177; Beier, 1963b: 145, fig. 147; Beier, 1970f: 320; Mahnert, 1977c: 83-84, figs 34a-e; Zaragoza, 1986: fig. 1; Harvey, 1991a: 378; Zaragoza, 2000a: 5-7, figs 1-3; Zaragoza, 2007a: 32.

Neobisium jeanneli (Ellingsen): Wolf, 1938: 611.

 

Type locality: Cueva del Pindal, Pimiango, Asturias, Spain.

Distribution: Spain (mainland).

Neobisium (Blothrus) karamani (Hadži)

Obisium (Blothrus) karamani Hadži, 1929: 61-71, figs 1-6.

Neobisium (Blothrus) karamani (Hadži): Beier, 1932a: 114; Roewer, 1937: 248; Beier, 1939d: 67; Beier, 1963b: 156, fig. 162; Beier, 1970f: 320; Ćurčić, 1974a: 20; Ćurčić, 1974b: 203-211, figs 13-35; Harvey, 1991a: 378.

Neobisium karamani (Hadži): Wolf, 1938: 612; Ćurčić, Dimitrijević and Legakis, 2004: 247-248.

 

Type locality: near Patiška, near Skopje, Skopje, Macedonia.

Distribution: Macedonia.

Neobisium (Blothrus) korabense Ćurčić

Neobisium korabense Ćurčić, 1982b: 146-149, figs 1a-e; Ćurčić, 1985b: 170-173, figs 1a-f, 2a-e; Ćurčić, Dimitrijević and Legakis, 2004: 248.

Neobisium (Blothrus) korabense Ćurčić: Harvey, 1991a: 378.

 

Type locality: Pećina Torbeški Most II Cave, near Ničpur, Mavrovi Anovi (as Mavrovi Hanovi), Mavrovi Anovi, Macedonia.

Distribution: Macedonia.

Neobisium (Blothrus) kosswigi Beier

Neobisium (Blothrus) kosswigi Beier, 1949: 5-6, fig. 5; Beier, 1965f: 90; Rack, 1971: 114; Beier, 1973e: 225; Mahnert, 1979b: 261; Harvey, 1991a: 378-379.

 

Type locality: Haci Akif Island, Beysehir Gölü, Anatolia, Konya, Turkey.

Distribution: Turkey.

Neobisium (Blothrus) kwartirnikovi Mahnert

Neobisium (Blothrus) kwartirnikovi Mahnert, 1972a: 383-388, figs 1-10, plate 1; Harvey, 1991a: 379; Petrov, 1997: 265.

 

Type locality: Cave Duhlata (as Duhlata Höhle), Bosnek, Pernik, Bulgaria.

Distribution: Bulgaria.

Neobisium (Blothrus) leruthi Beier

Neobisium (Blothrus) leruthi Beier, 1939c: 16-18, figs 11-12; Beier, 1963b: 154-155, fig. 160; Decou and Negrea, 1969: 523; Beier, 1970f: 320; Harvey, 1991a: 379; Ducháč, 1999b: 175-180, fig. 3; Ducháč, 2004c: 53, figs 5-7.

 

Type locality: Ghetarul de sub Zguràsti, Câmpeni, Turda, Romania.

Distribution: Romania.

Neobisium (Blothrus) lethaeum Beier

Neobisium (Blothrus) lethaeum Beier, 1938a: 7; Beier, 1970f: 320; Harvey, 1991a: 379; Ozimec, 2004: 387.

Neobisium lethaeum Beier: Ćurčić, 1988b: 47-48, figs 126-132; Ozimec, in Gottstein Matočec et al., 2002: 55 (as Neobisium letheum [sic]).

 

Type locality: Pećina Grabovica, near Grebci, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina; Croatia.

Neobisium (Blothrus) lethaeum acherusium Beier

Neobisium (Blothrus) lethaeum acherusium Beier, 1938a: 7; Beier, 1939d: 57, fig. 66; Beier, 1963b: 165; Ćurčić, 1974a: 20; Harvey, 1991a: 379.

 

Type locality: Ilijina Pećina, Trebinje, Federation of Bosnia and Herzegovina, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) lethaeum lethaeum Beier

Neobisium (Blothrus) lethaeum lethaeum Beier: Beier, 1939d: 55-56, fig. 64; Beier, 1963b: 165, fig. 171; Ćurčić, 1974a: 20

 

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) lethaeum parvum Beier

Neobisium (Blothrus) lethaeum parvum Beier, 1938a: 7; Beier, 1939d: 56-57, fig. 65; Beier, 1963b: 164; Ćurčić, 1974a: 21; Harvey, 1991a: 379; Ozimec, 2004: 387.

Neobisium lethaeum parvum Beier: Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 29.

 

Type locality: Jama na Maloj Žabi, (as 12 Meter tiefe Jama in der Mala Žaba Planina), NW. of Popovo Polje, Dubrovačko-Neretvanska, Croatia.

Distribution: Croatia.

Neobisium (Blothrus) lethaeum superbum Beier

Neobisium (Blothrus) lethaeum superbum Beier, 1938a: 7; Beier, 1939d: 58-59, fig. 68; Beier, 1963b: 164; Ćurčić, 1974a: 21; Harvey, 1991a: 379; Ozimec, 2004: 387.

Neobisium lethaeum superbum Beier: Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 58.

 

Type locality: Šipun Špilja, Cavtat, near Dubrovnik (as Ragusa), Dubrovačko-Neretvanska, Croatia.

Distribution: Croatia.

Neobisium (Blothrus) longidigitatum (Ellingsen)

Obisium (Roncus) longidigitatum Ellingsen, 1908a: 416-418; Ellingsen, 1912a: 169; Jeannel, 1926: 154.

Roncus (Roncus) longidigitatus (Ellingsen): Beier, 1932a: 129; Roewer, 1937: 249; Beier, 1963b: 186-187.

Roncus longidigitatus (Ellingsen): Wolf, 1938: 617.

Neobisium (Blothrus) longidigitatum (Ellingsen): Heurtault, 1975b: 631-635, figs 1-5; Heurtault, 1986c: 26 (as Neobisium (Blothrus) longedigitatum [sic]); Harvey, 1991a: 379; Delfosse, 2003: 35.

 

Type locality: Grotte d'Istaürdy, Pyrénées-Atlantiques (as Basses-Pyrénées), France.

Distribution: France (mainland).

Neobisium (Blothrus) maderi Beier

Neobisium (Blothrus) maderi Beier, 1938a: 8; Beier, 1939d: 66, fig. 79; Beier, 1963b: 162-163, fig. 169; Beier, 1970f: 320; Ćurčić, 1974a: 21; Harvey, 1991a: 380; Ozimec, 2004: 387.

Neobisium maderi Beier: Ćurčić, 1988b: 50-52, figs 138-163; Dimitrijević, 1997: 109-110, 111; Ozimec, in Gottstein Matočec et al., 2002: 55, fig. 41; Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 37.

 

Type locality: Ledenica na Podnožju Jabukovca, (as Ledenica Pećina), Mosor Planina, Splitsko-Dalmatinska, Croatia.

Distribution: Croatia.

Neobisium (Blothrus) maksimtodorovici Ćurčić and Dimitrijević

Neobisium maksimtodorovici Ćurčić, Dimitrijević and Mihajlova, 2002: 105-107, figs 1-8; Ćurčić, Dimitrijević and Legakis, 2004: 248.

 

Type locality: Momiček Cave, Belica, near Makedonski Brod, Dautica Mts, Makedonski Brod, Macedonia.

Distribution: Macedonia.

Neobisium (Blothrus) marcchagalli Ćurčić and S.B. Ćurčić

Neobisium marcchagalli Ćurčić and S.B. Ćurčić, in Ćurčić, Dimitrijević, S.B. Ćurčić, Tomić and N.B. Ćurčić, 2004: 93-97, figs 24-38; Ćurčić, Dimitrijević and Legakis, 2004: 249.

 

Type locality: Velja Peć Cave, Nikšićko Polje, Nikšić, Montenegro, Yugoslavia.

Distribution: Yugoslavia (Montenegro).

Neobisium (Blothrus) maxbeieri Dumitresco and Orghidan

Neobisium (Blothrus) beieri Dumitresco and Orghidan, 1970b: 101-104, figs 9-14 (junior primary homonym of Neobisium (Neobisium) beieri Verner, 1958).

Neobisium (Blothrus) maxbeieri Dumitresco and Orghidan, 1972: 247; Harvey, 1991a: 380 (replacement name for Neobisium (Blothrus) beieri Dumitresco and Orghidan).

 

Type locality: Grotte de Topolnita, near Drobeta-Turnu Severin, Mehedinti, Romania.

Distribution: Romania.

Neobisium (Blothrus) minutum (Tömösváry)

Blothrus minutus Tömösváry, 1882b: 235-237, plate 4 figs 11-13.

Obisium (Blothrus) brevipes J. Frivaldsky: Beier, 1928: 306 (in part).

Neobisium (Blothrus) minutum (Tömösváry): Beier, 1932a: 113-114, fig. 135; Roewer, 1937: 248; Beier, 1963b: 152-153, fig. 157; Rafalski, 1967: 12; Harvey, 1991a: 380; Ducháč, 2000: 7-10, fig. 2.

Neobisium (Blothrus) vulpinum Beier, 1936b: 85-86, fig. 1; Roewer, 1940: 343 (synonymised by Beier, 1963b: 153).

Neobisium minutum (Tömösváry): Verner, 1971: fig. 34.

 

Type locality: of Blothrus minutus: Mehádia, Caraş-Severin, Romania.

of Neobisium (Blothrus) vulpinum: "Schmiedeberg", Germany, but type material mislabelled (Beier, 1963b: 153).

Distribution: Hungary; Poland.

Neobisium (Blothrus) monasterii Mahnert

Neobisium (Blothrus) monasterii Mahnert, 1977c: 74-78, figs 20-24; Zaragoza, 1986: fig. 1; Harvey, 1991a: 380; Zaragoza, 2007a: 32.

 

Type locality: Cova de sa Campana, Escorca, Mallorca, Islas Baleares, Spain.

Distribution: Spain (Islas Baleares).

Neobisium (Blothrus) navaricum (Nonidez)

Obisium (Blothrus) navaricum Nonidez, 1925: 64-66, figs 5, 6a-b (as Obisium (Blothrus) navaricus [sic]); Jeannel, 1926: fig. 36.

Obisium navaricum Nonidez: Navás, 1925: 120.

Neobisium navaricum (Nonidez): Wolf, 1938: 613.

Neobisium (Blothrus) navaricum (Nonidez): Beier, 1932a: 108, fig. 125; Roewer, 1937: 248; Beier, 1939f: 175; Beier, 1963b: 138, fig. 138; Mahnert, 1977c: 73-74, figs 16-19; Estany, 1980b: 526, map 1; Zaragoza, 1986: fig. 1; Harvey, 1991a: 380; García Carrillo, 1998: 12; Zaragoza, 2007a: 32.

Neobisium (Blothrus) aff. navaricum (Nonidez): Zaragoza, 1986: fig. 1.

 

Type locality: Cueva de Malkorraundi, Gorriti, Navarra, Spain.

Distribution: Spain (mainland).

Neobisium (Blothrus) nonidezi (Bolivar)

Obisium nonidezi Bolivar, 1924: 101-103, fig. 1; Navás, 1925: 120.

Obisium (Blothrus) nonidezi Bolivar: Nonidez, 1925: 76-77, figs 14a-b.

Neobisium (Blothrus) nonidezi (Bolivar): Beier, 1932a: 117, fig. 142; Roewer, 1937: 248; Beier, 1939f: 176; Beier, 1963b: 141, fig. 143; Estany, 1980b: 527, map 2; Zaragoza, 1986: fig. 1; Harvey, 1991a: 380; Zaragoza, 2000a: 9-12, figs 9-13; Zaragoza, 2007a: 32-33.

Neobisium nonidezi (Bolivar): Wolf, 1938: 613.

 

Type locality: Cueva de Akelar, Lecumberri, Navarra, Spain.

Distribution: Spain (mainland).

Neobisium (Blothrus) occultum Beier

Neobisium (Blothrus) occultum Beier, 1938a: 7; Harvey, 1991a: 381.

 

Type locality: Lazarička Pećina, near Gacko Polje, Republika Srpska, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) occultum occultum Beier

Neobisium (Blothrus) occultum occultum Beier: Beier, 1939d: 60-61, figs 70-71; Beier, 1963b: 160, fig. 167; Beier, 1970f: 320; Ćurčić, 1974a: 21.

 

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) occultum sororium Beier

Neobisium (Blothrus) occultum sororium Beier, 1938a: 7; Beier, 1939d: 61, fig. 72; Beier, 1963b: 160-161; Ćurčić, 1974a: 21; Harvey, 1991a: 381.

 

Type locality: Mala Pečina, Miholjače, Gacko Polje, Republika Srpska, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) odysseum (Beier)

Obisium (Blothrus) odysseum Beier, 1929e: 451-453, figs 4, 5a, 6.

Neobisium (Blothrus) odysseum (Beier): Beier, 1932a: 109, fig. 127; Roewer, 1937: 248; Beier, 1939d: 59; Beier, 1963b: 159, fig. 165; Mahnert, 1975a: 179; Harvey, 1991a: 381.

Neobisium odysseum (Beier): Wolf, 1938: 613.

Not Neobisium (Blothrus) odysseum (Beier): Schawaller, 1985b: 5, 6, figs 8-11 (misidentification, see Neobisium (Ommatoblothrus) schawalleri Henderickx).

 

Type locality: Pantokrator above Barbati, Kérkira (as Korfu), Kérkira, Greece.

Distribution: Greece (mainland, Crete).

Neobisium (Blothrus) ohridanum Hadži

Neobisium (Blothrus) ohridanum Hadži, 1940: 129-135, figs 1a-c, 2, 3a-b, 4a-c, 5a-c; Beier, 1963b: 167-168; Ćurčić, 1974a: 21; Harvey, 1991a: 381; Ćurčić, Dimitrijević, Stojkovska, Stanović-Jovanović and Karamata, 1999: 9; Ćurčić, Dimitrijević and Legakis, 2004: 249-250.

 

Type locality: Mečkina Dupka, Zamne, near Ohrida, Ohrid, Macedonia.

Distribution: Macedonia.

Neobisium (Blothrus) peruni Ćurčić

Neobisium peruni Ćurčić, 1988b: 62-64, figs 261-272; Ozimec, in Gottstein Matočec et al., 2002: 55; Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 56.

Neobisium (Blothrus) peruni Ćurčić: Harvey, 1991a: 381; Ozimec, 2004: 387.

 

Type locality: Sonjina Jama, Lovka, Makarska, Biokovo Planina, Splitsko-Dalmatinska, Croatia.

Distribution: Croatia.

Neobisium (Blothrus) perunoides Ćurčić, Dimitrijević and Rađa

Neobisium perunoides Ćurčić, Dimitrijević and Rađa, 2006: 37-38, figs 1-8.

 

Type locality: Jama Na Vrhu Malog Šibenika Pothole, Mt Biokovo, Splitsko-Dalmatinska, Croatia.

Distribution: Croatia.

 

Neobisium (Blothrus) peyerimhoffi Heurtault

Neobisium (Blothrus) peyerimhoffi Heurtault, 1990c: 199-200, fig. 3.

 

Type locality: Ifri Bou Amane, Azerou Tidjer, Alger, Algeria.

Distribution: Algeria.

Neobisium (Blothrus) phineum Beier

Neobisium (Blothrus) phineum Beier, 1938a: 6; Harvey, 1991a: 381.

 

Type locality: Jama Plandište, Popovo Polje, Federation of Bosnia and Herzegovina, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) phineum extensum Beier

Neobisium (Blothrus) phineum extensum Beier, 1939d: 47, fig. 51; Beier, 1963b: 175; Ćurčić, 1974a: 21; Harvey, 1991a: 381.

 

Type locality: Neimenovana, Nevesinjskog Polje, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) phineum phineum Beier

Neobisium (Blothrus) phineum phineum Beier: Beier, 1939d: 46-47, fig. 50; Beier, 1963b: 170-171, fig. 178; Ćurčić, 1974a: 21.

 

Distribution: Bosnia-Herzegovina.

Neobisium (Blothrus) primitivum Beier

Neobisium (Blothrus) primitivum Beier, 1931a: 9-10, fig. 1; Beier, 1932a: 106, fig. 121; Roewer, 1937: 247; Beier, 1939f: 175; Estany, 1980b: 526, map 1; Harvey, 1991a: 381.

Neobisium primitivum Beier: Wolf, 1938: 613.

 

Type locality: Cueva de Mairuelegorreta, Monte Gorbea, Álava, País Vasco, Spain.

Distribution: Spain (mainland).

Neobisium (Blothrus) primitivum primaevum Beier

Neobisium (Blothrus) primaevum Beier, 1931a: 10-11, fig. 2; Beier, 1932a: 107, fig. 122; Roewer, 1937: 247; Beier, 1939f: 175.

Neobisium primaevum Beier: Wolf, 1938: 613.

Neobisium (Blothrus) primitivum primaevum Beier: Beier, 1963b: 137, fig. 136b; Zaragoza, 1986: fig. 1; Harvey, 1991a: 382; Zaragoza, 2007a: 33.

 

Type locality: Cueva de San Roque, Utzkorta, Vizcaya, País Vasco, Spain.

Distribution: Spain (mainland).

Neobisium (Blothrus) primitivum primitivum Beier

Neobisium (Blothrus) primitivum primitivum Beier: Beier, 1963b: 136-137, fig. 136a; Zaragoza, 1986: fig. 1; Zaragoza, 2007a: 33.

 

Distribution: Spain (mainland).

Neobisium (Blothrus) princeps Ćurčić

Neobisium (Blothrus) princeps Ćurčić, 1974b: 194-200, figs 1-12; Harvey, 1991a: 382; Ćurčić, Dimitrijević and Legakis, 2004: 250.

Neobisium (Blothrus) cf. princeps Ćurčić: Gardini, 1985a: 60-64, figs 17-23.

 

Type locality: Kalina Dupka Cave, 3 km N. of Lazaropolje, Macedonia.

Distribution: Greece (mainland) (doubtful); Macedonia.

Neobisium (Blothrus) pusillum Beier

Neobisium (Blothrus) pusillum Beier, 1939d: 41-42, fig. 43; Beier, 1963b: 167, fig. 173; Vachon and Gabbutt, 1964: figs 1, 12; Ćurčić, 1974a: 21; Harvey, 1991a: 382; Ozimec, 2004: 387.

 

Type locality: Črna Jama, near Planina, Slovenia.

Distribution: Croatia; Slovenia.

Neobisium (Blothrus) reimoseri (Beier)

Obisium spelaeum Schiödte: Valle, 1911: 24 (misidentification) (not seen).

Obisium (Blothrus) reimoseri Beier, 1929a: 361-364, figs 13, 14b; Beier, 1929e: fig. 5c.

Neobisium (Blothrus) reimoseri (Beier): Beier, 1932a: 109-110, fig. 128; Roewer, 1937: 248; Harvey, 1991a: 382; Gardini, 1995a: 4; Ozimec, 2004: 387.

Neobisium reimoseri (Beier): Wolf, 1938: 613.

Neobisium (Blothrus) reimoseri histricum Beier, 1939d: 42, fig. 44; Beier, 1963b: 169; Ćurčić, 1974a: 21 (synonymised by Mahnert, 1974b: 210).

Blothrus reimoseri (Beier): Caporiacco, 1949d: 139.

 

Type locality: of Obisium (Blothrus) reimoseri: Jama Draga, near Ponikve, Slovenia.

of Neobisium (Blothrus) reimoseri histricum: Jama Dimnice, near Slivje, Slovenia.

Distribution: Croatia; Italy (mainland); Slovenia.

Neobisium (Blothrus) reimoseri croaticum Beier

Neobisium (Blothrus) reimoseri croaticum Beier, 1939d: 43, fig. 45; Beier, 1963b: 169, fig. 175; Ćurčić, 1974a: 22; Harvey, 1991a: 382; Ozimec, 2004: 387.

Neobisium reimoseri croaticum Beier: Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 51.

 

Type locality: Pustinja Špilja, near Delnice, Primorsko-Goranska, Croatia.

Distribution: Croatia.

Neobisium (Blothrus) reimoseri reimoseri (Beier)

Neobisium (Blothrus) reimoseri reimoseri (Beier): Beier, 1939d: 42; Beier, 1963b: 168, fig. 174; Ćurčić, 1974a: 21; Mahnert, 1974b: 210-212, figs 7-10; Gardini, 1980c: 108, 128; Gardini, 2000: 71-72.

 

Distribution: Croatia; Italy (mainland); Slovenia.

Neobisium (Blothrus) remyi Beier

Neobisium (Blothrus) remyi Beier, 1939d: 67-68, fig. 81; Beier, 1963b: 156-157, fig. 163; Beier, 1970f: 320; Ćurčić, 1974a: 22; Harvey, 1991a: 382; Ćurčić, 1996: 344; Dimitrijević, 2000c: 39; Ćurčić, Dimitrijević and Legakis, 2004: 250.

 

Type locality: Županska Pećina, near Gorjo-Sela, Serbia, Yugoslavia.

Distribution: Yugoslavia (Montenegro, Serbia).

Neobisium (Blothrus) robustum (Nonidez)

Obisium (Blothrus) robustum Nonidez, 1925: 62-64, figs 3a-b, 4 (as Obisium (Blothrus) robustus [sic]).

Obisium robustum Nonidez: Navás, 1925: 120.

Neobisium (Blothrus) robustum (Nonidez): Beier, 1932a: 115, fig. 138; Roewer, 1937: 248; Beier, 1939f: 176; Beier, 1970f: 320; Estany, 1980b: 526, map 1; Bellés, 1987: fig. 41 (as Neobisium (Blothrus) robustus [sic]); Harvey, 1991a: 382-383.

Neobisium robustum (Nonidez): Wolf, 1938: 614.

 

Type locality: Cueva de San Adrián, Cegama, Guipúzcoa, País Vasco, Spain.

Distribution: Spain (mainland).

Neobisium (Blothrus) robustum escalerai Beier

Neobisium (Blothrus) escalerai Beier, 1931a: 14-15, fig. 6; Beier, 1932a: 115-116, fig. 139; Roewer, 1937: 248; Beier, 1939f: 176.

Neobisium escalerai Beier: Wolf, 1938: 611.

Neobisium (Blothrus) robustum escalerai Beier: Beier, 1963b: 140, fig. 141; Zaragoza, 1986: fig. 1; Harvey, 1991a: 383; Zaragoza, 2007a: 34.

 

Type locality: Cueva de Aitziquirri, Cegama, Guipúzcoa, País Vasco, Spain.

Distribution: Spain (mainland).

Neobisium (Blothrus) robustum robustum (Nonidez)

Neobisium (Blothrus) robustum robustum (Nonidez): Beier, 1963b: 139-140, fig. 140; Zaragoza, 1986: fig. 1; Zaragoza, 2007a: 33.

 

Distribution: Spain (mainland).

Neobisium (Blothrus) sbordonii Beier

Neobisium (Blothrus) sbordonii Beier, 1973e: 230-231, fig. 4; Harvey, 1991a: 383.

 

Type locality: Grotta Guezeu, Afsin, Kahramanmaraş, Turkey.

Distribution: Turkey.

Neobisium (Blothrus) slovacum Gulička

Neobisium (Blothrus) slovacum Gulička, 1977: 6-8, figs 1-4; Harvey, 1991a: 383; Ducháč, 1996: 153-156, figs 1-9; Ducháč, 1999b: 175-180, figs 1-2, 6-8; Ducháč and Mlejnek, 2000: 15-20, figs 1-2; Ducháč, 2004c: 52-53, figs 1-4; Ducháč, 2004e: 37 (not seen).

 

Type locality: Stará Brzotínská jaskyňa cave, near Ort Brzotín, Slovakia.

Distribution: Hungary; Slovakia.

Neobisium (Blothrus) spelaeum (Schiödte)

Blothrus spelaeus Schiödte, 1847: 80; Schiödte, 1851a: 23-26, plate 1 figs 2a-f; Schiödte, 1851b: 149, fig. 4; Hagen, 1870: 264; L. Koch, 1873: 41-42; G. Joseph, 1882: 21; Chamberlin, 1930: 11; Kästner, 1927b: fig. 11; Chamberlin, 1931a: 114, figs 32g-h, 41g-h.

Obisium troglodytes Schmidt, 1848: 10 (synonymised by Harvey, 1991a: 383).

Obisium spelaeum (Schiödte): Simon, 1881a: fig. 1.

Obisium (Blothrus) spelaeum (Schiödte): Daday, 1888: 128, 183 (as Obisium (Blothrus) spelaeus [sic]); Ellingsen, 1910a: 396; Beier, 1928: 306.

Neobisium (Blothrus) spelaeum (Schiödte): Beier, 1932a: 123-124, fig. 151; Roewer, 1937: 248; Beier, 1939d: 75; Beier, 1939e: 311; Beier, 1948b: 456; Beier, 1970f: 320; Harvey, 1991a: 383; Judson, 1997a: 41; Ozimec, 2004: 387.

Neobisium spelaeum (Schiödte): Wolf, 1938: 614; Ćurčić, 1988b: 52-54, figs 164-177; Ozimec, in Gottstein Matočec et al., 2002: 55.

Not Blothrus spelaeus Schiödte: Marchesetti, 1890: 159 (not seen) (misidentification; see Neobisium (Blothrus) spelaeum istriacum (G. Müller)); Roewer, 1931: 42 (misidentification; see Neobisium (Blothrus) spelaeum istriacum (G. Müller) and Neobisium (Blothrus) torrei (Simon)).

Not Obisium spelaeum (Schiödte): Hamann, 1896: 181 (not seen) (misidentification; see Neobisium (Blothrus) spelaeum istriacus (G. Müller)).

Not Obisium spelaeum Schiödte: Valle, 1911: 24 (not seen) (misidentification; see Neobisium (Blothrus) reimoseri (Beier)).

 

Type locality: of Blothrus spelaeus: Postojnska Jama (as Adelsberger Cave), near Postojna, Postojna, Slovenia.

of Obisium troglodytes: Postojnska Jama (as Adelsberger Cave), near Postojna, Postojna, Slovenia.

Distribution: Croatia; Italy (mainland); Slovenia.

Neobisium (Blothrus) spelaeum istriacum (G. Müller)

Blothrus spelaeus Schiödte: Marchesetti, 1890: 159 (not seen) (misidentification); Roewer, 1931: 42 (misidentification, in part).

Obisium spelaeum (Schiödte): Hamann, 1896: 181 (not seen) (misidentification).

Obisium (Blothrus) spelaeum istriacum G. Müller, 1931: 126-127.

Neobisium spelaeum istriacum (G. Müller): Wolf, 1938: 614.

Neobisium (Blothrus) spelaeum istriacum (G. Müller): Beier, 1939d: 75, fig. 90; Beier, 1963b: 178, fig. 188; Ćurčić, 1974a: 22; Gardini, 1980c: 108-109, 127, 128; Harvey, 1991a: 384; Gardini, 1995a: 4; Gardini, 2000: 72; Ozimec, 2004: 387.

 

Type locality: Jama Dimnice (as Grotta del Fumo), near Slivju, Istarska, Slovenia.

Distribution: Croatia; Italy (mainland); Slovenia.

Neobisium (Blothrus) spelaeum spelaeum (Schiödte)

Neobisium (Blothrus) spelaeum spelaeum (Schiödte): Beier, 1963b: 177-178, fig. 187; Ćurčić, 1974a: 22.

 

Distribution: Slovenia.

Neobisium (Blothrus) stankovici Ćurčić

Neobisium (Blothrus) stankovici Ćurčić, 1972e: 86-93, figs 1-19; Ćurčić, 1974a: 22; Harvey, 1991a: 384; Ćurčić, 1996: 344.

Neobisium stankovici Ćurčić: Ćurčić, 1976b: 176, fig. 2; Ćurčić, 1983b: fig. 2; Ćurčić, 1988e: fig. 1; Ćurčić, 1991b: 32-40, figs 1a-e, 2a-c, 3a-i, 4a-f, 5a-f; Dimitrijević, 2000c: 39; Ćurčić, Dimitrijević and Legakis, 2004: 250.

 

Type locality: Velika Pećina, Donja Držina, 7 km SE. of Pirot, Serbia, Yugoslavia.

Distribution: Yugoslavia (Serbia).

Neobisium (Blothrus) stitkovense Ćurčić and Dimitrijević

Neobisium stitkovense Ćurčić and Dimitrijević, in Ćurčić, Dimitrijević and S.B. Ćurčić, 2004: 479-482, figs 1-31; Ćurčić, Dimitrijević and Legakis, 2004: 250.

 

Type locality: Štitkovska Pećina, Štitkovo, near Nova Varoš, Mt Javor, Serbia, Yugoslavia.

Distribution: Yugoslavia (Serbia).

Neobisium (Blothrus) stygium Beier

Neobisium (Blothrus) stygium Beier, 1931a: 16-17, fig. 8; Beier, 1932a: 122-123, fig. 150; Caporiacco, 1937b: 41; Roewer, 1937: 248; Beier, 1970f: 320; Mahnert, 1974b: 212-216, figs 11-15; Helversen and Martens, 1972: 115-116, fig. 10b; Harvey, 1991a: 384; Ozimec, 2004: 387.

Neobisium (Blothrus) meuseli Beier, 1931a: 15-16, fig. 7; Beier, 1932a: 122, fig. 149; Roewer, 1937: 248 (synonymised by Mahnert, 1974b: 212).

Obisium (Blothrus) minoum Beier, 1931d: 95-96, figs 3c, 4 (synonymised by Helversen and Martens, 1972: 115).

Neobisium (Blothrus) minoum (Beier): Beier, 1932a: 121-122, fig. 148; Roewer, 1936: fig. 24a; Roewer, 1937: 248; Beier, 1939d: 72-73; Beier, 1963b: 175-176, fig. 184.

Neobisium (Blothrus) stygium padewiethi Beier, 1938a: 8; Beier, 1939d: 73, fig. 88; Beier, 1963b: 176-177; Ćurčić, 1974a: 22 (synonymised by Mahnert, 1974b: 212).

Neobisium meuseli Beier: Wolf, 1938: 612.

Neobisium minoum Beier: Wolf, 1938: 612.

Neobisium stygium Beier: Wolf, 1938: 615; Ćurčić, 1988b: 54-57, figs 178-220; Dimitrijević, 1997: 111; Ozimec, in Gottstein Matočec et al., 2002: 55.

Neobisium (Blothrus) stygium stygium Beier: Beier, 1939d: 73; Beier, 1963b: 176, fig. 185; Ćurčić, 1974a: 22.

Neobisium (Blothrus) stygium meuseli Beier: Beier, 1939d: 74; Beier, 1963b: 177; Ćurčić, 1974a: 22.

Neobisium (Blothrus) stygium csikii Beier, 1938a: 8; Beier, 1939d: 74, fig. 89; Beier, 1963b: 177, fig. 186; Ćurčić, 1974a: 22 (synonymised by Mahnert, 1974b: 212).

Neobisium stygium stygium Beier: Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 21.

Neobisium stygium meuseli Beier: Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 34.

Neobisium stygium csikii Beier: Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 63.

Neobisium stygium padewiethi Beier: Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 81.

 

 

Type locality: of Neobisium (Blothrus) stygium: Dumenčiča Špilja (as Höhle Duman), Karlovačka, Croatia.

of Neobisium (Blothrus) meuseli: Kosinski Špilja, Velebit Planina, Ličko-Senjska, Croatia.

of Obisium (Blothrus) minoum: Topolia, Crete, Greece, but quite possibly mislabelled (see Helversen and Martens, 1972).

of Neobisium (Blothrus) stygium padewiethi: Vlaška Peć, Kozica, Senj, Primorsko-Goranska, Croatia.

of Neobisium (Blothrus) stygium csikii: Špilja kod Lokava (as Risban- und Kormashöhle), Lokve, Gorski kotar, Goranska, Croatia.

Distribution: Croatia; Slovenia.

Neobisium (Blothrus) svetovidi Ćurčić

Neobisium svetovidi Ćurčić, 1988b: 60-62, figs 241-260; Ducháč, 2004d: 93-100, figs 1-8; Bedek, Gottstein Matočec, Jalžić, Ozimec and Štamol, 2006: 26.

Neobisium (Blothrus) svetovidi Ćurčić: Harvey, 1991a: 384; Ozimec, 2004: 387.

 

Type locality: Ivina Jama (as Ivina Jama Pothole), Krasno, Velebit Planina, Ličko-Senjska, Croatia.

Distribution: Croatia.

Neobisium (Blothrus) tantaleum Beier

Neobisium (Blothrus) tantaleum Beier, 1938a: 7; Beier, 1970f: 320; Harvey, 1991a: 385; Ozimec, 2004: 388.

Neobisium aff. tantaleum Beier: Ćurčić, 1988b: 57-60, figs 221-240; Dimitrijević, 1997: 111.

 

Type locality: Neimenovana Pećina, Vran, W. of Neretva River, Federation of Bosnia and Herzegovina, Bosnia-Herzegovina.

Distribution: Bosnia-Herzegovina; Croatia?.

Neobisium (Blothrus) tantaleum jablanicae Beier

Neobisium (Blothrus) tantaleum jablanicae Beier, 1938a: 8; Beier, 1939d: 64, fig. 77; Beier, 1963b: 166; Ćurčić, 1974a: 23; Harvey, 1991a: 385.