Chthonius (Chthonius) orthodactylus (Leach, 1817)

Chthonius (Chthonius) orthodactylus (Leach, 1817)

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Obisium orthodactylum Leach, 1817: 51, plate 141 fig. 2; Tulk, 1844: 55-57.

Obisium ischnocheles Hermann: Théis, 1832: 63-66, plate 1 figs 3, 3a-c (misidentification).

Chthonius orthodactylus (Leach): C.L. Koch, 1843: 79-80, fig. 808; Hagen, 1870: 268; L. Koch, 1873: 50-51; L. Koch, 1877: 181; Simon, 1879a: 73-74; Tömösváry, 1882b: 239-240, plate 5 figs 3-8; Becker, 1884: cclxiv; Pavesi, 1884: 457-458; Canestrini, 1885: no. 8, figs 1-4; Simon, 1885c: 50; Daday, 1888: 135, 191-192; Daday, 1889c: 25; Daday, 1889d: 81; O.P.-Cambridge, 1892: 209, plate A figs 7-9; Becker, 1896: 335; Simon, 1898d: 22 (misidentification, in part; see Paraliochthonius singularis (Menozzi)); Carpenter and Evans, 1899: 180-181; Simon, 1900b: 595; Evans, 1901a: 54; Carpenter, 1902: 220; Godfrey, 1908: 160; Ellingsen, 1909a: 211, 216; Warburton, 1909: 438; Ellingsen, 1910a: 401; Godfrey, 1910: 30; Jackson, 1910: 50; Kew, 1911a: 57, fig. 20; Kew, 1916a: 124; Daday, 1918: 2; Caporiacco, 1926: 75; Beier, 1928: 312-313; Kästner, 1928: 9; Schenkel, 1928: 71; Beier, 1929a: 365; Beier, 1929e: 454; Berland, 1929: 18; Bristowe, 1929a: 25; Chamberlin, 1929a: 71; Kew, 1930: 253-256, figs 1c-d; Vachon, 1934g: 163; Ionescu, 1936: 2; Roewer, 1937: 239; Savory, 1945: 180, 182; Caporiacco, 1950: 116; Caporiacco, 1951b: 103; Franciscolo, 1955: 125; Cerruti, 1959: 51 (as Chtonius [sic] orthodactylus); Baggini, 1961: 24; Bishop, 1967: 393; Marcuzzi, 1968: 267; Lazzeroni, 1970a: 207; Verner, 1971: fig. 4; Howes, 1972a: 108; Salmon, 1972: 67; Legg, 1973b: 430; Palmgren, 1973: 8; Legg, 1975a: 66; Legg, 1975b: 101, figs 1c, 4, 7c, 13a-b; Goddard, 1976b: 298-302, figs 2, 4; Goddard, 1979a: 100-104; Goddard, 1979b: 32; Judson, 1979a: 61; Matsuda, 1979: 201; Rundle, 1979: 47; Jones, 1980c: map 4; Schawaller, 1981b: 44, 50; Judson, 1987: 368; El-Hennawy, 1988b: 9; P.R. Harvey, 1992: 150; Fowles, 1994: 17; Legg and O'Connor, 1997: 109, fig. 2; Bell, Gates, Haughton, Macdonald, Smith, Wheater and Cullen, 1999: 237-239; Ducháč, 1999a: 14; Kárpáthegyi, 2007b: 91.

Chthonius (Chthonius) orthodactylus (Leach): Beier, 1932a: 50-51, figs 51, 56; Bristowe, 1938: 212; Beier, 1939e: 305; Cooreman, 1946b: 3; Beier, 1952e: 2; Beier, 1953e: 105; Beier and Franz, 1954: 453-454; G.O. Evans and Browning, 1954: 10, figs 9, 12; Beier, 1958c: 137; Beier, 1961f: 123; Beier, 1963b: 30, fig. 22; Beier, 1963a: 256; Beier, 1963h: 150; Lazzeroni, 1969a: 323; Lazzeroni, 1969b: 393-394; Lazzeroni, 1969c: 224; Lazzeroni, 1970c: 102; Legg, 1971a: 475, figs 1d, 2e; Ćurčić, 1974a: 12; Gardini, 1975: 2; Costantini, 1976: 122; Ćurčić, 1976b: 172; Crocker, 1978: 8; Leclerc and Heurtault, 1979: 239; Callaini, 1980a: 226; Gardini, 1980c: 100, 120, 121; Krumpál, 1980: 23; Inzaghi, 1981: 72; Krumpál and Kiefer, 1981: 128-129; Callaini, 1983d: 221-222; Bologna and Taglianti, 1985: 224, fig. 8; Callaini, 1986e: 236-237; Legg and Jones, 1988: 71-72, figs 14a, 14b(a-f); Ducháč, 1989a: 180; Gardini, 1991a: 172; Harvey, 1991a: 157; Ducháč, 1994: 140; Gardini, 1995a: 2; Platen, Blick, Bliss, Drogla, Malten, Martens, Scaher and Wunderlich, 1995: 54; Drogla and Blick, 1996: 37; Gardini, 2000: 26-28; Henderickx, 2000c: 67; Šťáhlavský, 2001: 80, fig. 2; Delfosse, 2003: 29; Mahnert, 2004: 463; Šťáhlavský and Král, 2004: 51, figs 1c, 5c; Ćurčić, Dimitrijević and Legakis, 2004: 235; Ozimec, 2004: 385; Tooren, 2005b: 92; Kárpáthegyi, 2007a: 82; Petrov and Šťáhlavský, 2007: 16; Šťáhlavský and Krásný, 2007: 430.

Not Obisium orthodactylum Leach: McIntire, 1868: 8 (misidentification; see Chthonius (Ephippiochthonius) tetrachelatus); McIntire, 1869: 247, fig. 219 (misidentification; see Chthonius (Ephippiochthonius) tetrachelatus).

Not Chthonius orthodactylus (Leach): Morley, 1930: 140-141 (misidentification; see Chthonius (Chthonius) ischnocheles (Hermann); Morley, 1940: 167 (misidentification; see Chthonius (Chthonius) ischnocheles (Hermann)).

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    Chthoniidae

    The Chthoniidae was first recognized as a distinct subfamily of pseudoscorpions by Daday (1888) and treated as a full family by Hansen (1894)....

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